Homologs of PP_4116 from Pseudomonas putida KT2440

isocitrate lyase
SEED: Isocitrate lyase (EC 4.1.3.1)
KEGG: isocitrate lyase

210 hits from BLASTp (E < 0.01)

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Orth Species Gene Name Description Fitness %Id Cov
  Pseudomonas putida KT2440 PP_4116 isocitrate lyase strong 100 100
o Pseudomonas sp. BP01 JOY50_RS06955 aceA isocitrate lyase insig. 98 100
o Pseudomonas sp. SVBP6 COO64_RS24260 aceA isocitrate lyase insig. 97 100
o Pseudomonas fluorescens GW456-L13 PfGW456L13_2633 Isocitrate lyase (EC 4.1.3.1) strong 96 100
o Pseudomonas protegens RBAN4 FABNJH_20150 aceA isocitrate lyase insig. 96 100
o Pseudomonas sp. DMC3 GFF1513 Isocitrate lyase strong 96 100
o Pseudomonas lactucae CFBP13502 GEMAOFIL_03641 Isocitrate lyase cofit 96 100
o Pseudomonas orientalis W4I3 QF045_RS18335 isocitrate lyase insig. 96 100
o Pseudomonas simiae WCS417 PS417_16570 isocitrate lyase strong 96 100
o Pseudomonas fluorescens FW300-N2E3 AO353_27735 isocitrate lyase strong 95 100
o Pseudomonas fluorescens SBW25 PFLU_RS18610 isocitrate lyase strong 95 100
o Pseudomonas fluorescens SBW25-INTG PFLU_RS18610 isocitrate lyase strong 95 100
o Pseudomonas fluorescens FW300-N1B4 Pf1N1B4_4042 Isocitrate lyase (EC 4.1.3.1) (from data) strong 95 100
o Pseudomonas sp. RS175 PFR28_02625 Isocitrate lyase strong 95 100
o Pseudomonas fluorescens FW300-N2C3 AO356_22080 isocitrate lyase strong 95 100
o Pseudomonas fluorescens FW300-N2E2 Pf6N2E2_279 Isocitrate lyase (EC 4.1.3.1) strong 95 100
o Pseudomonas syringae pv. syringae B728a Psyr_3196 isocitrate lyase strong 95 100
o Pseudomonas syringae pv. syringae B728a ΔmexB Psyr_3196 isocitrate lyase insig. 95 100
o Shewanella amazonensis SB2B Sama_2380 Isocitrate lyase (EC 4.1.3.1) (from data) strong 80 100
o Vibrio cholerae E7946 ATCC 55056 CSW01_03830 isocitrate lyase insig. 78 100
o Shewanella loihica PV-4 Shew_1276 Isocitrate lyase (EC 4.1.3.1) (from data) strong 78 99
o Shewanella sp. ANA-3 Shewana3_2942 isocitrate lyase (RefSeq) strong 78 99
o Shewanella oneidensis MR-1 SO1484 aceA Isocitrate lyase (EC 4.1.3.1) (from data) strong 78 99
o Variovorax sp. OAS795 ABID97_RS11115 isocitrate lyase strong 75 100
o Klebsiella pneumoniae MKP103 KDGMDA_10150 isocitrate lyase insig. 73 99
o Klebsiella pneumoniae MRSN742743 KFA93_RS21915 aceA;go_function=isocitrate isocitrate lyase insig. 73 99
o Paraburkholderia graminis OAS925 ABIE53_002536 isocitrate lyase strong 73 99
o Paraburkholderia bryophila 376MFSha3.1 H281DRAFT_00580 isocitrate lyase strong 73 99
o Escherichia coli ECRC100 OKFHMN_13265 aceA isocitrate lyase insig. 72 99
o Escherichia coli ECRC101 MCAODC_03765 aceA isocitrate lyase insig. 72 99
o Escherichia coli ECRC102 NIAGMN_11015 aceA isocitrate lyase insig. 72 99
o Escherichia coli ECRC98 JDDGAC_16890 aceA isocitrate lyase insig. 72 99
o Escherichia coli ECRC99 KEDOAH_14895 aceA isocitrate lyase no data 72 99
o Salmonella enterica subsp. enterica serovar Typhimurium str. MS1868 GFF1352 Isocitrate lyase (EC 4.1.3.1) insig. 72 99
o Serratia liquefaciens MT49 IAI46_23190 isocitrate lyase insig. 72 100
o Escherichia coli BL21 ECD_03887 isocitrate lyase insig. 72 99
o Escherichia coli ECOR27 NOLOHH_05410 aceA isocitrate lyase insig. 72 99
o Escherichia coli ECOR38 HEPCGN_11905 aceA isocitrate lyase insig. 72 99
o Escherichia coli ECRC62 BNILDI_06110 aceA isocitrate lyase insig. 72 99
o Escherichia coli HS(pFamp)R (ATCC 700891) OHPLBJKB_04035 Isocitrate lyase cofit 72 99
  Escherichia coli Nissle 1917 ECOLIN_RS23145 isocitrate lyase no data 72 99
  Escherichia coli Nissle 1917 ECOLIN_RS25430 isocitrate lyase no data 72 99
  Escherichia coli Nissle 1917 ECOLIN_RS25505 isocitrate lyase no data 72 99
  Escherichia coli Nissle 1917 ECOLIN_RS26870 isocitrate lyase no data 72 99
o Escherichia coli BW25113 b4015 aceA isocitrate lyase (NCBI) strong 72 99
o Rahnella sp. WP5 EX31_RS11745 isocitrate lyase insig. 71 100
o Dickeya dianthicola ME23 DZA65_RS20265 isocitrate lyase insig. 73 100
o Escherichia fergusonii Becca EFB2_04595 Isocitrate lyase insig. 72 99
o Enterobacter sp. TBS_079 MPMX20_00232 Isocitrate lyase strong 72 99
o Enterobacter asburiae PDN3 EX28DRAFT_4329 isocitrate lyase insig. 71 99
o Dickeya dadantii 3937 DDA3937_RS19055 isocitrate lyase insig. 72 100
o Dickeya dianthicola 67-19 HGI48_RS19145 isocitrate lyase insig. 72 100
o Castellaniella sp019104865 MT123 ABCV34_RS06915 isocitrate lyase insig. 71 99
o Dechlorosoma suillum PS Dsui_0011 isocitrate lyase strong 71 99
o Burkholderia phytofirmans PsJN BPHYT_RS12285 isocitrate lyase strong 72 99
o Herbaspirillum seropedicae SmR1 HSERO_RS10565 isocitrate lyase sig. 73 99
o Pectobacterium carotovorum WPP14 HER17_RS02295 isocitrate lyase insig. 71 100
o Variovorax sp. SCN45 GFF4829 Isocitrate lyase (EC 4.1.3.1) strong 71 100
o Klebsiella michiganensis M5al BWI76_RS01665 Isocitrate lyase (EC 4.1.3.1) (from data) strong 71 99
o Acidovorax sp. GW101-3H11 Ac3H11_2831 Isocitrate lyase (EC 4.1.3.1) (from data) strong 70 100
o Paraburkholderia sabiae LMG 24235 QEN71_RS06270 isocitrate lyase strong 71 99
o Hydrogenophaga sp. GW460-11-11-14-LB1 GFF4147 Isocitrate lyase (EC 4.1.3.1) strong 70 100
o Ralstonia solanacearum GMI1000 RS_RS06810 isocitrate lyase insig. 71 99
o Cupriavidus basilensis FW507-4G11 RR42_RS12200 isocitrate lyase insig. 70 99
o Ralstonia solanacearum IBSBF1503 RALBFv3_RS00150 isocitrate lyase insig. 71 99
o Ralstonia solanacearum UW163 UW163_RS13480 isocitrate lyase insig. 71 99
o Ralstonia solanacearum PSI07 RPSI07_RS17335 isocitrate lyase insig. 70 99
o Dyella japonica UNC79MFTsu3.2 ABZR86_RS13490 isocitrate lyase insig. 71 97
o Pantoea agglomerans CFBP13505 P0401 PagCFBP13516_RS18250 aceA isocitrate lyase cofit 69 99
o Rhodanobacter sp000427505 FW510-R12 LRK53_RS03375 isocitrate lyase insig. 71 97
o Rhodanobacter sp. FW510-T8 OKGIIK_13320 aceA isocitrate lyase cofit 71 97
o Lysobacter sp. OAE881 ABIE51_RS00190 isocitrate lyase cofit 70 99
o Ralstonia sp. UNC404CL21Col ABZR87_RS11590 isocitrate lyase strong 69 99
o Pantoea sp. MT58 IAI47_18005 isocitrate lyase insig. 68 99
o Rhodanobacter denitrificans MT42 LRK55_RS10225 isocitrate lyase insig. 71 97
o Rhodanobacter denitrificans FW104-10B01 LRK54_RS10585 isocitrate lyase strong 71 97
o Xanthomonas campestris pv. campestris strain 8004 Xcc-8004.311.1 Isocitrate lyase (EC 4.1.3.1) insig. 69 98
o Mycobacterium tuberculosis H37Rv Rv0467 Isocitrate lyase Icl (isocitrase) (isocitratase) no data 63 98
o Pontibacter actiniarum KMM 6156, DSM 19842 CA264_09255 isocitrate lyase insig. 61 100
o Agrobacterium fabrum C58 Atu0607 isocitrate lyase strong 63 95
o Sinorhizobium meliloti 1021 SMc00768 isocitrate lyase strong 61 95
o Rhizobium sp. OAE497 ABIE40_RS03255 isocitrate lyase strong 62 95
o Azospirillum sp. SherDot2 MPMX19_05830 Isocitrate lyase insig. 60 95
o Azospirillum brasilense Sp245 AZOBR_RS02405 isocitrate lyase strong 60 95
o Brevundimonas sp. GW460-12-10-14-LB2 A4249_RS02595 isocitrate lyase strong 60 95
  Mycobacterium tuberculosis H37Rv Rv1915 Probable isocitrate lyase AceAa [first part] (isocitrase) (isocitratase) (Icl) insig. 37 59
  Caulobacter crescentus NA1000 CCNA_01841 isocitrate lyase sig. 29 81
  Caulobacter crescentus NA1000 Δfur CCNA_01841 isocitrate lyase cofit 29 81
  Acinetobacter radioresistens SK82 MPMX26_02319 Isocitrate lyase strong 30 74
  Alteromonas macleodii MIT1002 MIT1002_02831 Isocitrate lyase strong 28 90
  Pseudomonas stutzeri RCH2 Psest_2014 Isocitrate lyase (EC 4.1.3.1) (from data) strong 29 83
  Sphingobium sp. HT1-2 GFF2635 Isocitrate lyase (EC 4.1.3.1) cofit 28 96
  Rhodopseudomonas palustris CGA009 TX73_022775 isocitrate lyase no data 27 95
  Pseudomonas aeruginosa MRSN321 DY961_RS04045 isocitrate lyase strong 29 83
  Pseudomonas aeruginosa PA14 IKLFDK_13225 isocitrate lyase strong 29 83
  Pseudomonas aeruginosa PUPa3 DQ20_RS35075 isocitrate lyase strong 29 83
  Pseudomonas sp. S08-1 OH686_00170 Isocitrate lyase strong 28 83
  Acinetobacter baumannii LAC-4 RR41_RS08965 isocitrate lyase insig. 29 74
  Sphingomonas koreensis DSMZ 15582 Ga0059261_1963 Isocitrate lyase insig. 26 93
  Cupriavidus basilensis FW507-4G11 RR42_RS25845 isocitrate lyase sig. 28 80
  Ralstonia sp. UNC404CL21Col ABZR87_RS08330 isocitrate lyase sig. 28 83
  Marinobacter adhaerens HP15 HP15_3234 isocitrate lyase strong 28 80
  Pseudomonas segetis P6 ACVTMO_RS12700 isocitrate lyase strong 27 80
  Mycobacterium tuberculosis H37Rv Rv1916 Probable isocitrate lyase AceAb [second part] (isocitrase) (isocitratase) (Icl) insig. 51 22
  Hydrogenophaga sp. GW460-11-11-14-LB1 GFF4149 Isocitrate lyase (EC 4.1.3.1) insig. 70 16
  Lysobacter sp. OAE881 ABIE51_RS06015 methylisocitrate lyase sig. 29 61
  Rhodanobacter sp. FW510-T8 OKGIIK_06565 prpB methylisocitrate lyase strong 29 59
  Escherichia coli Nissle 1917 ECOLIN_RS02150 methylisocitrate lyase insig. 29 63
  Escherichia fergusonii Becca EFB2_03706 2-methylisocitrate lyase insig. 29 63
  Escherichia coli ECRC100 OKFHMN_08900 prpB methylisocitrate lyase insig. 29 63
  Escherichia coli ECRC101 MCAODC_28130 prpB methylisocitrate lyase insig. 29 63
  Escherichia coli ECRC102 NIAGMN_06955 prpB methylisocitrate lyase cofit 29 63
  Escherichia coli ECRC98 JDDGAC_12555 prpB methylisocitrate lyase insig. 29 63
  Escherichia coli ECRC99 KEDOAH_19065 prpB methylisocitrate lyase no data 29 63
  Escherichia coli BL21 ECD_00286 2-methylisocitrate lyase insig. 29 63
  Escherichia coli ECOR27 NOLOHH_01775 prpB methylisocitrate lyase insig. 29 63
  Escherichia coli ECOR38 HEPCGN_07485 prpB methylisocitrate lyase insig. 29 63
  Escherichia coli ECRC62 BNILDI_23190 prpB methylisocitrate lyase cofit 29 63
  Escherichia coli BW25113 b0331 prpB 2-methylisocitrate lyase (NCBI) sig. 29 63
  Rhodanobacter sp000427505 FW510-R12 LRK53_RS09460 methylisocitrate lyase sig. 29 59
  Escherichia coli HS(pFamp)R (ATCC 700891) OHPLBJKB_03339 2-methylisocitrate lyase insig. 29 63
  Rhodanobacter denitrificans MT42 LRK55_RS00470 methylisocitrate lyase strong 29 59
  Rhodanobacter denitrificans FW104-10B01 LRK54_RS00695 methylisocitrate lyase strong 29 59
  Ralstonia solanacearum GMI1000 RS_RS17740 methylisocitrate lyase insig. 30 59
  Ralstonia sp. UNC404CL21Col ABZR87_RS16745 methylisocitrate lyase strong 30 59
  Pseudomonas sp. S08-1 OH686_01690 methylisocitrate lyase strong 31 59
  Ralstonia solanacearum PSI07 RPSI07_RS00400 methylisocitrate lyase insig. 29 59
  Ralstonia solanacearum IBSBF1503 RALBFv3_RS22820 methylisocitrate lyase insig. 29 59
  Ralstonia solanacearum UW163 UW163_RS23440 methylisocitrate lyase insig. 29 59
  Herbaspirillum seropedicae SmR1 HSERO_RS15660 2-methylisocitrate lyase no data 31 58
  Ralstonia solanacearum UW163 UW163_RS11055 methylisocitrate lyase insig. 30 64
  Ralstonia solanacearum IBSBF1503 RALBFv3_RS02670 methylisocitrate lyase insig. 29 64
  Methylophilus sp. DMC18 GFF469 2-methylisocitrate lyase no data 29 59
  Ralstonia solanacearum PSI07 RPSI07_RS14645 methylisocitrate lyase insig. 29 64
  Salmonella enterica subsp. enterica serovar Typhimurium str. MS1868 GFF4263 Methylisocitrate lyase (EC 4.1.3.30) insig. 29 63
  Ralstonia solanacearum GMI1000 RS_RS10050 methylisocitrate lyase insig. 29 64
  Xanthomonas campestris pv. campestris strain 8004 Xcc-8004.3981.1 Methylisocitrate lyase (EC 4.1.3.30) strong 30 59
  Pseudomonas segetis P6 ACVTMO_RS13040 prpB methylisocitrate lyase insig. 28 59
  Alteromonas macleodii MIT1002 MIT1002_03129 Methylisocitrate lyase strong 31 54
  Pseudomonas sp. DMC3 GFF5230 2-methylisocitrate lyase cofit 28 59
  Pseudomonas fluorescens GW456-L13 PfGW456L13_4118 Methylisocitrate lyase (EC 4.1.3.30) strong 34 41
  Vibrio cholerae E7946 ATCC 55056 CSW01_06735 methylisocitrate lyase cofit 29 59
  Ralstonia sp. UNC404CL21Col ABZR87_RS14590 methylisocitrate lyase insig. 29 63
  Cupriavidus basilensis FW507-4G11 RR42_RS11260 Methylisocitrate lyase (EC 4.1.3.30) (from data) strong 28 60
  Pseudomonas stutzeri RCH2 Psest_2321 methylisocitrate lyase strong 33 41
  Pseudomonas aeruginosa MRSN321 DY961_RS08515 prpB;go_function=methylisocitrate methylisocitrate lyase strong 29 59
  Pseudomonas aeruginosa PA14 IKLFDK_03445 methylisocitrate lyase strong 29 59
  Pseudomonas aeruginosa PUPa3 DQ20_RS44335 prpB;go_function=methylisocitrate methylisocitrate lyase strong 29 59
  Acinetobacter baumannii LAC-4 RR41_RS18485 methylisocitrate lyase insig. 27 65
  Paraburkholderia sabiae LMG 24235 QEN71_RS17555 methylisocitrate lyase insig. 28 59
  Pseudomonas orientalis W4I3 QF045_RS22905 methylisocitrate lyase insig. 28 63
  Pseudomonas protegens RBAN4 FABNJH_09625 prpB methylisocitrate lyase insig. 27 59
  Kangiella aquimarina DSM 16071 B158DRAFT_1343 methylisocitrate lyase strong 31 46
  Pseudomonas fluorescens FW300-N2E3 AO353_00900 2-methylisocitrate lyase strong 27 59
  Pseudomonas fluorescens SBW25 PFLU_RS22690 methylisocitrate lyase insig. 33 40
  Pseudomonas fluorescens SBW25-INTG PFLU_RS22690 methylisocitrate lyase strong 33 40
  Pseudomonas simiae WCS417 PS417_21080 2-methylisocitrate lyase no data 33 40
  Paraburkholderia sabiae LMG 24235 QEN71_RS31070 methylisocitrate lyase insig. 32 55
  Pseudomonas lactucae CFBP13502 GEMAOFIL_04361 2-methylisocitrate lyase cofit 33 40
  Pseudomonas sp. RS175 PFR28_02928 2-methylisocitrate lyase strong 32 41
  Pseudomonas fluorescens FW300-N1B4 Pf1N1B4_3819 Methylisocitrate lyase (EC 4.1.3.30) sig. 32 41
  Pseudomonas fluorescens FW300-N2C3 AO356_20865 2-methylisocitrate lyase strong 32 41
  Pseudomonas sp. SVBP6 COO64_RS09900 prpB methylisocitrate lyase insig. 32 41
  Pseudomonas fluorescens FW300-N2E2 Pf6N2E2_6061 Methylisocitrate lyase (EC 4.1.3.30) (from data) strong 32 41
  Shewanella loihica PV-4 Shew_1820 prpB 2-methylisocitrate lyase (RefSeq) strong 30 55
  Acinetobacter radioresistens SK82 MPMX26_01289 2-methylisocitrate lyase no data 35 39
  Pseudomonas sp. BP01 JOY50_RS25750 prpB methylisocitrate lyase insig. 32 41
  Marinobacter adhaerens HP15 HP15_1932 methylisocitrate lyase 2 no data 29 60
  Dyella japonica UNC79MFTsu3.2 ABZR86_RS08480 methylisocitrate lyase strong 29 59
  Shewanella oneidensis MR-1 SO0345 prpB methylisocitrate lyase (NCBI ptt file) strong 29 55
  Pseudomonas putida KT2440 PP_2334 2-methylisocitrate lyase strong 32 41
  Pseudomonas syringae pv. syringae B728a Psyr_2085 methylisocitrate lyase insig. 32 40
  Pseudomonas syringae pv. syringae B728a ΔmexB Psyr_2085 methylisocitrate lyase insig. 32 40
  Shewanella sp. ANA-3 Shewana3_3825 prpB 2-methylisocitrate lyase (RefSeq) strong 29 57
  Shewanella amazonensis SB2B Sama_3294 prpB 2-methylisocitrate lyase (RefSeq) strong 30 52
  Variovorax sp. SCN45 GFF1663 Isocitrate lyase (EC 4.1.3.1) strong 33 42
  Castellaniella sp019104865 MT123 ABCV34_RS01670 methylisocitrate lyase cofit 27 59
  Xanthobacter sp. DMC5 GFF2763 2,3-dimethylmalate lyase insig. 43 20
  Acidovorax sp. GW101-3H11 Ac3H11_2326 Methylisocitrate lyase (EC 4.1.3.30) insig. 32 35
  Dinoroseobacter shibae DFL-12 Dshi_0689 isocitrate lyase family protein (RefSeq) sig. 46 20
  Azospirillum brasilense Sp245 AZOBR_RS24315 carboxyvinyl-carboxyphosphonate phosphorylmutase insig. 33 35
  Variovorax sp. SCN45 GFF268 Carboxyvinyl-carboxyphosphonate phosphorylmutase (EC 2.7.8.23) strong 43 19
  Azospirillum brasilense Sp245 AZOBR_RS27785 carboxyvinyl-carboxyphosphonate phosphorylmutase insig. 32 35
  Variovorax sp. OAS795 ABID97_RS19370 isocitrate lyase/PEP mutase family protein insig. 24 74
  Burkholderia phytofirmans PsJN BPHYT_RS25670 carboxyvinyl-carboxyphosphonate phosphorylmutase insig. 33 31
  Paraburkholderia bryophila 376MFSha3.1 H281DRAFT_02542 2-Methylisocitrate lyase, PEP mutase family insig. 31 40
  Rhodospirillum rubrum S1H Rru_A2320 Ankyrin (NCBI) insig. 31 41
  Paraburkholderia bryophila 376MFSha3.1 H281DRAFT_00986 2-Methylisocitrate lyase, PEP mutase family insig. 31 31
  Azospirillum sp. SherDot2 MPMX19_04657 2,3-dimethylmalate lyase insig. 31 37
  Variovorax sp. SCN45 GFF1681 Methylisocitrate lyase (EC 4.1.3.30) insig. 31 37
  Cupriavidus basilensis FW507-4G11 RR42_RS22175 carboxyvinyl-carboxyphosphonate phosphorylmutase insig. 29 51
  Cupriavidus basilensis FW507-4G11 RR42_RS22765 carboxyvinyl-carboxyphosphonate phosphorylmutase insig. 31 37
  Paraburkholderia bryophila 376MFSha3.1 H281DRAFT_06209 carboxyvinyl-carboxyphosphonate phosphorylmutase insig. 33 36
  Hydrogenophaga sp. GW460-11-11-14-LB1 GFF3032 Methylisocitrate lyase (EC 4.1.3.30) no data 42 20
  Rhodopseudomonas palustris CGA009 TX73_012370 methylisocitrate lyase insig. 32 32
  Azospirillum brasilense Sp245 AZOBR_RS27820 carboxyvinyl-carboxyphosphonate phosphorylmutase insig. 42 19
  Dinoroseobacter shibae DFL-12 Dshi_1092 carboxyvinyl-carboxyphosphonatephosphorylmutase (RefSeq) insig. 39 19
  Burkholderia phytofirmans PsJN BPHYT_RS26855 carboxyvinyl-carboxyphosphonate phosphorylmutase sig. 32 35
  Xanthobacter sp. DMC5 GFF2758 2,3-dimethylmalate lyase insig. 40 19
  Hydrogenophaga sp. GW460-11-11-14-LB1 GFF5012 putative methylisocitrate lyase insig. 38 19
  Variovorax sp. OAS795 ABID97_RS18835 oxaloacetate decarboxylase insig. 39 19
  Bosea sp. OAE506 ABIE41_RS16605 isocitrate lyase/PEP mutase family protein insig. 29 36
  Paraburkholderia graminis OAS925 ABIE53_005902 2-methylisocitrate lyase-like PEP mutase family enzyme strong 30 35
  Paraburkholderia bryophila 376MFSha3.1 H281DRAFT_02892 carboxyvinyl-carboxyphosphonate phosphorylmutase insig. 39 19
  Azospirillum sp. SherDot2 MPMX19_02797 2,3-dimethylmalate lyase insig. 29 34
  Hydrogenophaga sp. GW460-11-11-14-LB1 GFF1578 Methylisocitrate lyase (EC 4.1.3.30) insig. 38 19
  Rhizobium sp. OAE497 ABIE40_RS01965 isocitrate lyase/phosphoenolpyruvate mutase family protein insig. 36 25
  Pseudomonas sp. BP01 JOY50_RS14465 oxaloacetate decarboxylase insig. 39 19
  Caulobacter crescentus NA1000 CCNA_01980 PEP phosphonomutase related enzyme insig. 39 19
  Caulobacter crescentus NA1000 Δfur CCNA_01980 PEP phosphonomutase related enzyme cofit 39 19