Pairwise Alignments

Query, 4328 a.a., Pyoverdine chromophore precursor synthetase PvdL from Pseudomonas fluorescens FW300-N2E2

Subject, 4161 a.a., non-ribosomal peptide synthetase from Pseudomonas fluorescens FW300-N2C3

 Score = 2744 bits (7114), Expect = 0.0
 Identities = 1610/3725 (43%), Positives = 2166/3725 (58%), Gaps = 163/3725 (4%)

Query: 668  EALPQSLAQNRLWITWRLDPTSHAYNIPGALHLRGELDEDALRTSFQQLIERHESLRTRF 727
            + L  S AQ R W  W+LDP S AY++P AL LRG LD  AL  SF  L+ RHESLRT F
Sbjct: 48   DTLSLSFAQERQWFLWQLDPESAAYHMPTALRLRGPLDIVALEQSFTALVARHESLRTTF 107

Query: 728  LERDGVALQQIDPAGEFKLHVIDLGDLPADSREARARQVREDEAATPFDLEKGPLLRVTL 787
                   +QQ+  A   +L    L  LP    E + RQ+ E E    FDL++GPLLR  L
Sbjct: 108  QTHGEQVVQQVHAAAPVRLQADTLEQLPD---EMQLRQLIEGETRQLFDLQQGPLLRTRL 164

Query: 788  LRLDEDDHQLLVTLHHIIADGWSMSVLIEEFSRLYAAASQGQVAELAPLPLQYADYGSWQ 847
            LRL  DDH L+VTLHHI+ADGWSM V++EE    YAA SQG+  +L  LP+QY DY  WQ
Sbjct: 165  LRLAADDHVLIVTLHHIVADGWSMQVMVEELVDGYAAFSQGRTPQLPALPIQYTDYAIWQ 224

Query: 848  RQWLAQGEAERQLSYWKQQLGAEQSALVLGTDHPRSAQRQRSAARHSLRLDKDLGEALRH 907
            R W+  GE ERQL YW  +LG +Q  L L TD PR A      AR  L +D  L   L+H
Sbjct: 225  RHWMEAGERERQLEYWTARLGGQQPMLELPTDFPRPAVMSYRGARLELAIDAALAGDLKH 284

Query: 908  VAQAHDATPFMLLLAAFQALLQRYTGQSDIRIGVPNANRPRLETQGLIGFFINTQVLRGQ 967
            +AQ    + FMLLLA+FQALL RY+GQ+DIR+GVP ANR R+E + LIGFF+NTQVL   
Sbjct: 285  MAQREGVSLFMLLLASFQALLHRYSGQNDIRVGVPTANRNRVEAERLIGFFVNTQVLDSD 344

Query: 968  VDSRQPFAALLAQAREATLGAQAHQDLPFEQLLEAIPEAREQG---LFQVMFNHQQRDLS 1024
            V S+  F ALL Q ++  LGAQAHQ+LPFEQL+EA+   R      LFQVMFNH+ +  +
Sbjct: 345  VQSQDSFGALLQQVKQTALGAQAHQELPFEQLVEALQPERNLSHSPLFQVMFNHRTQGPA 404

Query: 1025 ALRRLPGLLAEELPWHSREAKFDLQLHSEEDRNGRLTLSFDYADELFEAATIERLATHYG 1084
                +  L  EEL W S  A+FDL L + E   G L+    YA +LF A TI R+A H+ 
Sbjct: 405  GEGPVGALKVEELRWDSHTAQFDLTLETVEHSEG-LSAGLIYATDLFGAETIARMAGHWQ 463

Query: 1085 NLLRAVCAGPQQAIGDVPLLTADEHSQQQ---QWSAAPCAPASQWLPELLNEQARNTPQR 1141
            NLLRA+ A P Q IG++PLL A E   Q    Q +  P AP + ++ +L   QA  TP  
Sbjct: 464  NLLRAIVADPAQRIGELPLLDAAERQAQLDNGQRTPTP-APEAGFVHQLFERQAARTPDA 522

Query: 1142 IALRWDGGQLDYAELHAQANRLAHYLRDKGVGPDVCVAIAAERSPQLLIGVLAIIKAGGA 1201
             AL  DG +  Y +L+A+AN+LA  L+  GVGPDV V IA ERS  +L+G+LAI+KAGGA
Sbjct: 523  PALVIDGERWSYGQLNARANQLAGKLQSLGVGPDVLVGIAVERSAHMLVGLLAILKAGGA 582

Query: 1202 YVPLDADYPAERLAYMLDDSGVDLLLTQTALLERL------------------------- 1236
            Y+PLD  +P +RLAYM++DSG++LLLTQ++LL++L                         
Sbjct: 583  YLPLDPAFPQDRLAYMIEDSGIELLLTQSSLLDQLRLSCGSELARDSGGSVNIDVGGEGL 642

Query: 1237 ------PACDGVSVIAM----DALHLEQWPSNPPGLHLHGDHLAYVIYTSGSTGQPKGVG 1286
                  P   GV V  +    +   L+ +    P   L  +HLAY IYTSGSTG+PKGV 
Sbjct: 643  IASKLAPTVFGVGVTTLVLDQEGDWLDGYACEAPAPRLDAEHLAYAIYTSGSTGKPKGVM 702

Query: 1287 NTHAALAERLQWMQATYGLDAIDVLMQKAPISFDVSVWECFWPLITGSQLVLAGPGEHRD 1346
              H AL   +  M    G+ A D  +     SFD+   E + PL+ G+ +VL G    +D
Sbjct: 703  VRHGALTNFVASMAQAPGMTAQDRALSLTTFSFDIFGLEIYVPLMVGACIVLTGQHVAQD 762

Query: 1347 PHRIAQLVQQFGVTTLHFVPPLLSLFVDEPLSARCTSLRRLFSGGEALPAELRNRVLAQL 1406
            P+ +  L+    V  +   P    + +D   +      + L  GGEALP EL  R+LA  
Sbjct: 763  PYAVLALIAAEQVNVVQATPSTWRMLLDNEHAQGLRGCKCL-CGGEALPQELAVRMLAL- 820

Query: 1407 PSVQLHNRYGPTETAINVTHW---HCTEADGERSPIGRPLGNVLCRVLDSDLNPVPAGVP 1463
               ++ N YGPTET I    W   H  +    +  +G P+ N    ++ +DL PVP GV 
Sbjct: 821  -GGEVWNLYGPTETTI----WSALHPLQTQAAQPWLGGPIDNTALYIVGADLMPVPYGVA 875

Query: 1464 GELCIGGLGLARGYLGRPGLSAERFVADPLGQAGARLYRTGDRARWNADGVIEYLGRLDQ 1523
            GEL IGG GLARGY  RP L+AERFV  P   AG RLYRTGD AR  ADG++EYLGR+D 
Sbjct: 876  GELLIGGQGLARGYFQRPALTAERFVPHPF-VAGERLYRTGDLARQRADGILEYLGRIDH 934

Query: 1524 QVKLRGFRVEPQEIEARLLAQDGVAQVAVLVRNTSAGPQLIGYYTAPEV-----AQGQDE 1578
            QVK+RGFR+E  EIEARLL QD V + AVL +  + G  L+ Y           A  Q  
Sbjct: 935  QVKIRGFRIELGEIEARLLEQDSVREAAVLAQEAAGGQALVAYIVPTAELNDLDAGAQGA 994

Query: 1579 LNTRLKAALAAELPDYMVPAQLLRLDVMPLSPSGKLDRRALPEP---QWQVREHVEPASV 1635
            L   L A L   LPDYMVPA ++ L   PL+P+GKLDR+ LP+P   QWQ + +V P + 
Sbjct: 995  LRDSLTARLKVNLPDYMVPAHMVVLPRFPLTPNGKLDRKQLPKPDASQWQ-KTYVAPQTP 1053

Query: 1636 LEQQIAGIWRQVLGLARIGLRDDFFALGGHSLLATQIISRTRQTCDVELPLRTLFEASEL 1695
            ++Q +A IW  +L + ++GL D FF LGGHSLLATQ++SR RQ  ++++PLR LF  + L
Sbjct: 1054 VQQAVATIWADILKVPQVGLDDSFFELGGHSLLATQVMSRIRQQLNIQVPLRLLFAHNSL 1113

Query: 1696 GAFAEQVRLIQASGQANRQPPIEKVDRSQPVPLSYSQQRMWFLWQMEPDSPAYNVGGMAR 1755
             AF E +     +  A+ +P I ++ R QP+ LS++QQR WFLWQ+EP S AY++    R
Sbjct: 1114 RAFVEAM----GTASAHDEPGIPRIPRDQPLALSFAQQRQWFLWQLEPTSSAYHIPAALR 1169

Query: 1756 LRGVLDVGRFEAALQALILRHETLRTTFPSVDGVAHQQVHGETGLRMDWKDFSMLAFDVR 1815
            LRG LD    + +  AL+ RHE+LRT F   +    Q +     L +  +    L   V 
Sbjct: 1170 LRGALDRAALQQSFDALVARHESLRTRFVLDNERPLQVIDEPAPLALAVES---LEAGVD 1226

Query: 1816 EQRVQQLADSEAHKPFDLETGPLLRACLVKTAEHEHYFVLTLHHIVTEGWAMDIFARELS 1875
            E+ ++ L ++E  + FDL+ GPLLR   ++    +H  +LT HHIV++ W+M I   +L 
Sbjct: 1227 ERTLRGLIEAETRQLFDLQRGPLLRVKWLQLGAEDHVLILTQHHIVSDAWSMQIMVDDLL 1286

Query: 1876 ALYEAFIDDRESPLEPLPVQYLDYSVWQRQWLESGERQRQLDYWTTQLGREHPLLELPGD 1935
             LY  F   +   L PLP+QY DY  WQRQWL +GE+QRQLDYW  QLG E P+LELP D
Sbjct: 1287 QLYAGFSQGQPVALAPLPIQYADYGHWQRQWLAAGEQQRQLDYWIEQLGGEQPILELPTD 1346

Query: 1936 RPRPSVQSHRGELFRFDLSDDLAARVRAFNAQQGLTLFMTMTAALAVLLYRYSGQTDLRI 1995
             PRP+ QS+RG      +   LA  ++    Q   TLFM + A    LL+RYSGQ D+R+
Sbjct: 1347 HPRPAQQSYRGARLPIVIEPALAEGLKRLAQQHNSTLFMLLLAGFQTLLHRYSGQDDIRV 1406

Query: 1996 GAPVANRIRPESEGLIGAFLNTQVLRCQLDGQMSVGELFEQVRHTVIEGQSHQDLPFDHL 2055
            G P+ANR R E+E L+G F+NTQVL+  +DG +   +L +QV+   +  Q+HQDLPF+ L
Sbjct: 1407 GVPIANRNRVETERLLGFFVNTQVLKADIDGNLRFDQLLDQVKQRALGAQAHQDLPFEQL 1466

Query: 2056 VEALQPPRSAAYNPLFQVMCNVQRWEFQQSRTLAGMTVEYLVNDARATKFDLNLEVTDLD 2115
            VE L+P R+ ++ PLFQVM N      +    L G+++E L  D+   +FDL L+  +  
Sbjct: 1467 VERLEPARNLSHTPLFQVMFNHASDARRDLSALPGLSLEQLTWDSGTAQFDLTLDTFEYA 1526

Query: 2116 QRLGCCLTYSTDLFDEPRIARMAGHWRNLLEALLDDPARRLSELPLLDTREQQQLLDSLG 2175
              L   LTY++DLF+   IAR+  HW NLL A++ +P +R++ELP+LD  +++ +++   
Sbjct: 1527 DGLSAALTYASDLFEPSTIARLGEHWLNLLRAIVANPTQRIAELPMLDRTQRRLIVEDWN 1586

Query: 2176 VEPGEHHLDRCLHELFDEQARMRPDAPALTFAGQTLTYAQLDSSANRLAWMLRERGVGPQ 2235
                ++     +H L + Q    P A AL F GQ+L+YAQL++ AN+LA  L ERGVGP 
Sbjct: 1587 RTYADYPNHLPVHHLIEAQVERDPHAEALVFEGQSLSYAQLNARANQLAHALIERGVGPD 1646

Query: 2236 VRVGLALERSLEMVIGLLAILKAGGAYVPLDPEYPLDRLHYMIEDSGISLLLS------- 2288
            V VG++++RS+EMV+GLLAILKAGGAYVPLDPEYP +RL YMIEDSGI LLL+       
Sbjct: 1647 VLVGISVQRSVEMVVGLLAILKAGGAYVPLDPEYPPERLAYMIEDSGIELLLTCGSELAR 1706

Query: 2289 ------------DRAMLQALGALPDGVGCWCLEDDSPALAQYPDHALPLINLPQHQAYLI 2336
                        +  +   L     G+    L+ D   L+ +     P   + Q+ AY+I
Sbjct: 1707 DSGGSVNIDVEGEGLIASKLAPTGGGLNLLVLDGDDDWLSDHSTDNPPARAVAQNLAYVI 1766

Query: 2337 YTSGSTGKPKGVVVSHGEIAMHCQAVIQRFGMRPDDCELHFYSINFDAATERLLVPLLSG 2396
            YTSGSTGKPKG   SHG +      + Q +G+   D  L     +FD +      PL++G
Sbjct: 1767 YTSGSTGKPKGAGNSHGALTNRLCWMQQAYGLEAGDSVLQKTPFSFDVSVWEFFWPLMTG 1826

Query: 2397 AHVVLRAQGQW-DAEEICGLIRQHRINILGFTPSYGSQLAQWLATQGQTLPVRMCITGGE 2455
              +V+ A G   D +++  LI +  I  L F PS   Q     A   +   +   +  GE
Sbjct: 1827 VRLVVAAPGDHRDPQKLVSLIERESITTLHFVPSM-LQAFVLDANVSRCTCLTRIVCSGE 1885

Query: 2456 ALTGEHLQRIRAAFNPSLFFNAYGPTETVVMPLASLAPEQLEEGAASVPIGSVIGARMAY 2515
            AL  +  Q++ A    +  +N YGPTE  +           +EG  +VPIG  I   M Y
Sbjct: 1886 ALPVDAQQQVFAKLPNARLYNLYGPTEAAIDVTHWTC---RDEGRDAVPIGEPIANLMTY 1942

Query: 2516 ILDADLALVPPGATGELYVGGAGLAQGYHQRPGMTAERFVADPFATDGGRLYRTGDLVRQ 2575
            ILD +L  VP G  GELY+ G GLA+GYH+RP +TAERFVA PF   G RLYRTGDL R 
Sbjct: 1943 ILDDELNPVPSGVNGELYLSGQGLARGYHRRPELTAERFVASPFVA-GERLYRTGDLARY 2001

Query: 2576 RADGLVEYLGRIDHQVKIRGFRIELGEIETRLLEHDAVREAVVLALDTPGGKQLAGYLVS 2635
            R DG+++Y+GRIDHQVK+RGFRIELGEIE RLLEH AVREAVV+A D+  GK L+GY+V+
Sbjct: 2002 RPDGVIDYVGRIDHQVKVRGFRIELGEIEARLLEHAAVREAVVVAQDSRSGKVLSGYVVA 2061

Query: 2636 DVASQDDGRQAALRDALKNHLKAQLPDYMVPTHLILLASMPLTANGKLDRRALPMPDPEL 2695
                  D     LR+AL  HL+  LP++MVPTHL  L  MPL+ NGKLDR+ALP PD   
Sbjct: 2062 QAGEASD--WPLLREALLAHLRLNLPEHMVPTHLTRLEHMPLSPNGKLDRKALPAPDLAE 2119

Query: 2696 NRQHYVAPSNALEQTLAAIWCDVLNVTQVGLNDNFFELGGDSILSIQVVSRARQQGIHFT 2755
             ++ + AP NA+E+ L  IW  VL V +VG  DNFF LGGDSI SIQVVSRAR QGI  T
Sbjct: 2120 RQEAFTAPVNAIERALVDIWQGVLGVERVGTADNFFALGGDSINSIQVVSRARAQGIGLT 2179

Query: 2756 PRDLFQHQTVQTLAAVATRSQQVLAEQGLLQGESGLTPIQHWFFERPIPNRHHWNQALVL 2815
            P+DLF HQ +Q LA VAT + + + +QG + G+S L P+QHWFF   IP+R HWNQ+++L
Sbjct: 2180 PKDLFLHQNIQALARVATAADEGVIDQGPVTGDSLLVPMQHWFFASDIPHRDHWNQSVLL 2239

Query: 2816 EPTTALAPQPLEQALRSLLQHHDALRLGFDEST-GHWQAEHQPLSDAALLRHVSVDTAQD 2874
             P   L    L+QAL+ ++ HHDALRL F ++  G W A HQ +    L   +   TAQD
Sbjct: 2240 TPREPLDIAHLQQALQHVVAHHDALRLRFTQANDGRWHASHQAVDKQPL--EIWQRTAQD 2297

Query: 2875 CEALFA---ETQSRFNLATGPLLRAVLAQFPDGQQRLLIVIHHLVVDGVSWRVLMDDLQT 2931
               + A   + Q   +L  GPLLR  L    DG  RL + IHHLV+DGVSWRVL++DLQ 
Sbjct: 2298 AAEIIAIADQAQRSLDLQHGPLLRCSLIAVQDGSSRLHLAIHHLVMDGVSWRVLLEDLQN 2357

Query: 2932 VYRPLAIGQAAQLSAKTSPMRDWAARLQAYAGSESLREELAWWQAQLSGAQVPLPCANPE 2991
            VYR L   QA  L  K++  + WAA+LQ YA SE+L +EL +W  QL     PLP A PE
Sbjct: 2358 VYRQLRANQAPALPGKSTAFKAWAAQLQDYAHSETLNKELDYWVGQLDQPLPPLPRARPE 2417

Query: 2992 GGLLERHAETVSVRLDAERTRQLLQQAPSAYRTQVNDLLLTALARVLCRWSGHASTLVQL 3051
            G     H  +VS RLDA +T++LL+QAP+AYRTQVNDLLL+ALA+V+C+W+G  + LVQL
Sbjct: 2418 GSQAAAHGRSVSTRLDATQTQRLLKQAPAAYRTQVNDLLLSALAQVVCQWTGQPACLVQL 2477

Query: 3052 EGHGREALFDDIDLTRTVGWFTSVYPVRLSPATEGFGASIKAIKEQLRAVPHKGLGHGVL 3111
            EGHGRE LF+ IDL+RTVGWFTS++PV L+PA +    SIK IKEQLRAVP+KGLG+GVL
Sbjct: 2478 EGHGREELFEGIDLSRTVGWFTSLFPVLLTPAGDP-AESIKRIKEQLRAVPNKGLGYGVL 2536

Query: 3112 RYLADAATRQTMANLPHAAITFNYLGQLDQTLGHDALFQPLDAPLGAIHDPDAPLPNELS 3171
            RYL   A +Q +A LP A ITFNYLGQ DQ+    ALF P +   GA H  D  L N L+
Sbjct: 2537 RYLGAEAVQQRLAKLPPARITFNYLGQFDQSFDEQALFVPSEEGSGAGHGDDVALGNWLT 2596

Query: 3172 VDSQVSGGELLLRWTFSRERHDRQAIATLAGAYLDELQRLIEHCLTDEAGGLTPSDFPLA 3231
            +D ++  G+L L WTFS +  D   +  LA AY   LQ+LI+HC TD   GLTPSDFPLA
Sbjct: 2597 IDGRIFNGQLTLDWTFSGDVFDEATLQRLADAYAVALQQLIDHCCTDGHLGLTPSDFPLA 2656

Query: 3232 KLTQGQLDSLPVPAGQIEDVYPLTPMQEGMLLHTLLEPGTGLYYMQDRYRINSELDPQRF 3291
            +L+Q QL+ L VP  Q+ED+YPL+PMQ+GML HTL +  +GLY  Q    ++  LD +RF
Sbjct: 2657 RLSQAQLNGLTVPVAQVEDIYPLSPMQDGMLFHTLADDSSGLYINQISLPVHG-LDTERF 2715

Query: 3292 AQAWQAVIARHEALRASFCWNVG-EDMLQIIHRPGSTPVDYLDWSEVPEDAQEAKLQALL 3350
            A+AW +VI R + LR SF W  G    +QI+HR    P+   DW    +D  E  + A  
Sbjct: 2716 ARAWTSVIEREDILRTSFHWQDGLSAPVQIVHRHVELPLRIEDWR--GQDISEQAISARA 2773

Query: 3351 KNEREAGFDLLNQAPFHLRLIRVGAARYWFMMSNHHILIDAWCRSLLMDDFFEIYTALGE 3410
              +  AGFDL       + L+R+   RY  + + HHIL+D W  S L  +  + Y     
Sbjct: 2774 TADYLAGFDLSRAPLQRVLLLRLEDDRYQMIWTCHHILMDGWSSSRLFGEVLQFY---AH 2830

Query: 3411 NREPQLTVPPRYRDYIGWLQHQSLAEARQWWQRNLQGFERTTPIPGDRPFLREHAGDSGG 3470
               P      RYR++I WLQ Q      ++W+  L   E+ T +   +     H     G
Sbjct: 2831 GHVPSRN--GRYREFIEWLQRQDQGALERFWRSRLDTLEQATSL--SQAMFPRHVEALPG 2886

Query: 3471 MIVGDCYTRLDARDGARLRELAQAHQLTVNTFAQAAWALVLRRVSGDRDVLFGVTVAGRP 3530
                  Y+R D    ARL++  +A Q+T NT  Q AW L+L+R +G   V+FG TVAGRP
Sbjct: 2887 HEA--LYSRWDVAQTARLQQACRALQITPNTLIQGAWLLLLQRFTGQNTVVFGATVAGRP 2944

Query: 3531 VELPQMQRTVGLFINSIALRVQMPEDHQRCSVRQWLSDLLDHNMQLREYEYLPLVSIQEA 3590
              L      +GLFIN++ + +Q  +  Q   + QWL  L  +N+ LRE+ + PL  IQ  
Sbjct: 2945 AGLVNADNILGLFINTLPV-IQTLDPEQ--PLDQWLQQLQAYNLDLREHSHAPLADIQRW 3001

Query: 3591 SELPKGQPLFDSLFVFENAPVEVSVLDRAQSLNATSDSGRTH--TNFPLTAVCYPGDDLG 3648
            S L  GQ LFDS+ VFEN P++  + D+ Q    +    R H  TNFP+      G+ L 
Sbjct: 3002 SGL-GGQGLFDSIIVFENYPIDERLGDQ-QDTGLSFGESRNHDVTNFPMDLAVNLGETLS 3059

Query: 3649 LHLSYDQRYFDEATVQGLLSEFKRLLLALMEGLHGDMSELPLIGAQEQDFLIEGCNQSEH 3708
            +   + +  F    V+ +    +  L A+++  H  +  L  +  ++     +       
Sbjct: 3060 IEYLFLRNAFSVEAVERIRRTLEDTLEAMIDAPHVCLGNLQRLSLEQWQAFAQWSAAPNG 3119

Query: 3709 AYPLEQSYVELFEARVAAHPHRPAVSCLDTSYSYAELNARSNRLGHALIAAGVGLDQPVA 3768
             Y   Q   EL      + P   A+ C D S  Y EL  R+NRL H LI  G G +  V 
Sbjct: 3120 CYQ-AQLLPELIGRHAQSRPQATALVCGDDSLDYGELERRANRLAHRLIELGAGPEVVVG 3178

Query: 3769 LLAERDAELLGMIVGSFKAGAGYLPLDPGLPSQRLGRIIELSRTPLLVCTQACQAQ---- 3824
            +  ER  +++  ++   K+G+ Y+PLD   P +RL  +IE S   LL+     QA+    
Sbjct: 3179 VALERSVDMVVALLAVMKSGSAYVPLDIDYPPERLAFMIEDSAMALLLTHSRVQARLPDL 3238

Query: 3825 ---AQALLDEFACSGRPKLLVWEAVQASALSLENPGVYSGPDNLAYVIYTSGSTGLPKGV 3881
               A+  +D F   G   L              + G+  G  NLAY+IYTSGSTG PKGV
Sbjct: 3239 DGLARLAIDGFDSRGYSDL------------PPHSGLLGG--NLAYLIYTSGSTGRPKGV 3284

Query: 3882 MVEQRGMLNNQLSKVPYLALSEVDVIAQTASQSFDISVWQFLAAPLFGARVDIVPNAIAH 3941
             V    M  +  +      + E        S +FD +  ++L+  LF     ++ +    
Sbjct: 3285 AVAHGPMSMHCQAIAGLYDMDEQTRELHFMSFAFDGAHERWLST-LFSGGTLVIRDGALW 3343

Query: 3942 DPQGLLEHVQQQGITVLESVPSLIQGM---LAQERIGLDGLRWMLPTGEAMPPELAHQWL 3998
             P+   + + + GIT+    P+ ++ +    A   +    +R     G+A+P +   Q  
Sbjct: 3344 TPEQTFDALHRHGITIACFPPAYLKQLAEYAANSSLAPPLVRVYCFGGDAVPDQTFEQVK 3403

Query: 3999 LRYPDIGLVNAYGPAECSDDVAFFRVDLAS-TRGTYLPIGTPTDNNRLYLLDGALDLVPL 4057
                     N YGP E       ++V  +      Y PIG       LY+LD  L+ +P 
Sbjct: 3404 AALRPQFFTNGYGPTETVVTPMLWKVPASERCEAAYAPIGRAVGERALYVLDDNLNPLPP 3463

Query: 4058 GAVGELCVAGIGVGRGYVSDPLRTAPVFVPNPFGAPGERLYRTGDLARRRSDGVLEYVGR 4117
            G  GEL + G G+ RGY   P  +A  FVP+PFG PG+RLYR+GDL R+R DGVL+YVGR
Sbjct: 3464 GIAGELYIGGAGIARGYHRRPDLSAERFVPDPFGQPGDRLYRSGDLVRQRVDGVLDYVGR 3523

Query: 4118 IDHQVKIRGYRIELGEIEARLHEQPEVRDGAVGVQEGVNGKHLVGYLVATDSALKPAERL 4177
            IDHQ+K+RG+RIELGE+EA L +  +V D  V  ++  +GK L+GY+V  D         
Sbjct: 3524 IDHQIKVRGFRIELGEVEASLRQLDDVSDALVIARDSASGKQLIGYVVTPDGVAIG---- 3579

Query: 4178 ERIKQRLRAELPEYMVPLHWLWLERLPLNANGKLDRKALPALEIGQLQSQDYQAPGNELE 4237
            +R+K  LRA+LP+YMVP   + LE LP+  NGKLDR+ALP  E    +  +Y    N  E
Sbjct: 3580 DRLKAELRAQLPDYMVPAQVICLEALPVTPNGKLDRQALPEPE---FKGAEYVPGRNRDE 3636

Query: 4238 QTLADIWAQVLKVERVGVRDNFFELGGHSLLATQIASRVQK--ALQRNVPLRAMFECSTV 4295
            Q LA++WA+VL+VERVG+ DNFFELGG S+L+ Q+ SRV+    L+  + LR +    T+
Sbjct: 3637 QLLAEVWAEVLQVERVGITDNFFELGGDSILSLQVVSRVRNHGELKMELKLRDLMRGQTI 3696

Query: 4296 QELAD 4300
            + + D
Sbjct: 3697 EAIFD 3701



 Score = 1133 bits (2930), Expect = 0.0
 Identities = 819/2251 (36%), Positives = 1151/2251 (51%), Gaps = 241/2251 (10%)

Query: 16   QRRAALTPDRVALRFLADTPGQGVVLSYRDLDLRARTIAAALQAEAALGDRAV-LLFPSG 74
            +R+AA TPD  AL         G   SY  L+ RA  +A  LQ+     D  V +     
Sbjct: 513  ERQAARTPDAPALVI------DGERWSYGQLNARANQLAGKLQSLGVGPDVLVGIAVERS 566

Query: 75   PDYVAAFFGCLYAGVIAVPAYPPESTRRHHQERLLSIIADAEPRLLLTRSDLREPLQ--- 131
               +      L AG   +P  P        Q+RL  +I D+   LLLT+S L + L+   
Sbjct: 567  AHMLVGLLAILKAGGAYLPLDPA-----FPQDRLAYMIEDSGIELLLTQSSLLDQLRLSC 621

Query: 132  ----------------------AIESAPPV--LCVDTLDPSRAGQWR--------MPTLQ 159
                                  A + AP V  + V TL   + G W          P L 
Sbjct: 622  GSELARDSGGSVNIDVGGEGLIASKLAPTVFGVGVTTLVLDQEGDWLDGYACEAPAPRLD 681

Query: 160  DDDIAFLQYTSGSTALPKGVQVSHGNLVANELLIRHGFGIDLNPDDVIVSWLPLYHDM-G 218
             + +A+  YTSGST  PKGV V HG L      +    G  +   D  +S      D+ G
Sbjct: 682  AEHLAYAIYTSGSTGKPKGVMVRHGALTNFVASMAQAPG--MTAQDRALSLTTFSFDIFG 739

Query: 219  LIGGLLQPVFSGVPCVLMSPAYFLARPLRWLEAISEYGGTISGGPDFAYRLCSERVSDSA 278
            L   +  P+  G  C++++  +    P   L  I+     +       +R+  +      
Sbjct: 740  L--EIYVPLMVGA-CIVLTGQHVAQDPYAVLALIAAEQVNVVQATPSTWRMLLDNEHAQG 796

Query: 279  LQRLDLSGWRVAYSGSEPIRLDTLERFAEKFAPCGFTPDNFMASYGLAEATLFVAGTPRR 338
            L+            G E +     +  A +    G    N    YG  E T++ A  P  
Sbjct: 797  LRGCK------CLCGGEALP----QELAVRMLALGGEVWNL---YGPTETTIWSALHP-- 841

Query: 339  HGIPSLRVDDAALAQNRAEPGQGSAVMSCGVSQPGHAVLIVEPATAQPLADNQVGEVWAA 398
                        L    A+P  G  +         +  L +  A   P+     GE+   
Sbjct: 842  ------------LQTQAAQPWLGGPI--------DNTALYIVGADLMPVPYGVAGELLIG 881

Query: 399  GPSIALGYWRNPEASAKTFVQHD---GRTWLRTGDLGFLR-DGELFITGRLKDLLIVRGH 454
            G  +A GY++ P  +A+ FV H    G    RTGDL   R DG L   GR+   + +RG 
Sbjct: 882  GQGLARGYFQRPALTAERFVPHPFVAGERLYRTGDLARQRADGILEYLGRIDHQVKIRGF 941

Query: 455  NLYPQDIEQAIEREVEVVRKGRVAAFAVNDGGEEGIGIAAEISRSVQKILPPEALIKAIR 514
             +   +IE  +  E + VR+  V A         G  + A I  + +          A+R
Sbjct: 942  RIELGEIEARL-LEQDSVREAAVLA----QEAAGGQALVAYIVPTAELNDLDAGAQGALR 996

Query: 515  QAVAEACQ-QAPSVVALLNPGALPK---TSSGKLQRSACRSRLADGSLDSYAQFPSAEVE 570
             ++    +   P  +   +   LP+   T +GKL R                Q P  +  
Sbjct: 997  DSLTARLKVNLPDYMVPAHMVVLPRFPLTPNGKLDRK---------------QLPKPDAS 1041

Query: 571  DADES--APSSELQTLIGQIWREQLQVEQVQADDHFFLLGGNSIAATQMIARLREILGLE 628
               ++  AP + +Q  +  IW + L+V QV  DD FF LGG+S+ ATQ+++R+R+ L ++
Sbjct: 1042 QWQKTYVAPQTPVQQAVATIWADILKVPQVGLDDSFFELGGHSLLATQVMSRIRQQLNIQ 1101

Query: 629  LNLRLLFEAPTLAAFAEAVANQQQDGGQPQGAISALSRDEALPQSLAQNRLWITWRLDPT 688
            + LRLLF   +L AF EA+          +  I  + RD+ L  S AQ R W  W+L+PT
Sbjct: 1102 VPLRLLFAHNSLRAFVEAMGTAS---AHDEPGIPRIPRDQPLALSFAQQRQWFLWQLEPT 1158

Query: 689  SHAYNIPGALHLRGELDEDALRTSFQQLIERHESLRTRFLERDGVALQQIDPAGEFKLHV 748
            S AY+IP AL LRG LD  AL+ SF  L+ RHESLRTRF+  +   LQ ID      L V
Sbjct: 1159 SSAYHIPAALRLRGALDRAALQQSFDALVARHESLRTRFVLDNERPLQVIDEPAPLALAV 1218

Query: 749  IDLGDLPADSREARARQVREDEAATPFDLEKGPLLRVTLLRLDEDDHQLLVTLHHIIADG 808
                 L A   E   R + E E    FDL++GPLLRV  L+L  +DH L++T HHI++D 
Sbjct: 1219 ---ESLEAGVDERTLRGLIEAETRQLFDLQRGPLLRVKWLQLGAEDHVLILTQHHIVSDA 1275

Query: 809  WSMSVLIEEFSRLYAAASQGQVAELAPLPLQYADYGSWQRQWLAQGEAERQLSYWKQQLG 868
            WSM +++++  +LYA  SQGQ   LAPLP+QYADYG WQRQWLA GE +RQL YW +QLG
Sbjct: 1276 WSMQIMVDDLLQLYAGFSQGQPVALAPLPIQYADYGHWQRQWLAAGEQQRQLDYWIEQLG 1335

Query: 869  AEQSALVLGTDHPRSAQRQRSAARHSLRLDKDLGEALRHVAQAHDATPFMLLLAAFQALL 928
             EQ  L L TDHPR AQ+    AR  + ++  L E L+ +AQ H++T FMLLLA FQ LL
Sbjct: 1336 GEQPILELPTDHPRPAQQSYRGARLPIVIEPALAEGLKRLAQQHNSTLFMLLLAGFQTLL 1395

Query: 929  QRYTGQSDIRIGVPNANRPRLETQGLIGFFINTQVLRGQVDSRQPFAALLAQAREATLGA 988
             RY+GQ DIR+GVP ANR R+ET+ L+GFF+NTQVL+  +D    F  LL Q ++  LGA
Sbjct: 1396 HRYSGQDDIRVGVPIANRNRVETERLLGFFVNTQVLKADIDGNLRFDQLLDQVKQRALGA 1455

Query: 989  QAHQDLPFEQLLEAIPEAREQG---LFQVMFNH---QQRDLSALRRLPGLLAEELPWHSR 1042
            QAHQDLPFEQL+E +  AR      LFQVMFNH    +RDLSA   LPGL  E+L W S 
Sbjct: 1456 QAHQDLPFEQLVERLEPARNLSHTPLFQVMFNHASDARRDLSA---LPGLSLEQLTWDSG 1512

Query: 1043 EAKFDLQLHSEEDRNGRLTLSFDYADELFEAATIERLATHYGNLLRAVCAGPQQAIGDVP 1102
             A+FDL L + E  +G L+ +  YA +LFE +TI RL  H+ NLLRA+ A P Q I ++P
Sbjct: 1513 TAQFDLTLDTFEYADG-LSAALTYASDLFEPSTIARLGEHWLNLLRAIVANPTQRIAELP 1571

Query: 1103 LLTADEHSQ-QQQWSAAPCAPASQWLP--ELLNEQARNTPQRIALRWDGGQLDYAELHAQ 1159
            +L   +     + W+    A     LP   L+  Q    P   AL ++G  L YA+L+A+
Sbjct: 1572 MLDRTQRRLIVEDWNRT-YADYPNHLPVHHLIEAQVERDPHAEALVFEGQSLSYAQLNAR 1630

Query: 1160 ANRLAHYLRDKGVGPDVCVAIAAERSPQLLIGVLAIIKAGGAYVPLDADYPAERLAYMLD 1219
            AN+LAH L ++GVGPDV V I+ +RS ++++G+LAI+KAGGAYVPLD +YP ERLAYM++
Sbjct: 1631 ANQLAHALIERGVGPDVLVGISVQRSVEMVVGLLAILKAGGAYVPLDPEYPPERLAYMIE 1690

Query: 1220 DSGVDLLLTQTALLER--------------------LPACDGVSVIAMDALHLEQWPS-- 1257
            DSG++LLLT  + L R                     P   G++++ +D    + W S  
Sbjct: 1691 DSGIELLLTCGSELARDSGGSVNIDVEGEGLIASKLAPTGGGLNLLVLDG--DDDWLSDH 1748

Query: 1258 ---NPPGLHLHGDHLAYVIYTSGSTGQPKGVGNTHAALAERLQWMQATYGLDAIDVLMQK 1314
               NPP   +   +LAYVIYTSGSTG+PKG GN+H AL  RL WMQ  YGL+A D ++QK
Sbjct: 1749 STDNPPARAV-AQNLAYVIYTSGSTGKPKGAGNSHGALTNRLCWMQQAYGLEAGDSVLQK 1807

Query: 1315 APISFDVSVWECFWPLITGSQLVLAGPGEHRDPHRIAQLVQQFGVTTLHFVPPLLSLFVD 1374
             P SFDVSVWE FWPL+TG +LV+A PG+HRDP ++  L+++  +TTLHFVP +L  FV 
Sbjct: 1808 TPFSFDVSVWEFFWPLMTGVRLVVAAPGDHRDPQKLVSLIERESITTLHFVPSMLQAFVL 1867

Query: 1375 EPLSARCTSLRRLFSGGEALPAELRNRVLAQLPSVQLHNRYGPTETAINVTHWHCTEADG 1434
            +   +RCT L R+   GEALP + + +V A+LP+ +L+N YGPTE AI+VTHW C +   
Sbjct: 1868 DANVSRCTCLTRIVCSGEALPVDAQQQVFAKLPNARLYNLYGPTEAAIDVTHWTCRDEGR 1927

Query: 1435 ERSPIGRPLGNVLCRVLDSDLNPVPAGVPGELCIGGLGLARGYLGRPGLSAERFVADPLG 1494
            +  PIG P+ N++  +LD +LNPVP+GV GEL + G GLARGY  RP L+AERFVA P  
Sbjct: 1928 DAVPIGEPIANLMTYILDDELNPVPSGVNGELYLSGQGLARGYHRRPELTAERFVASPF- 1986

Query: 1495 QAGARLYRTGDRARWNADGVIEYLGRLDQQVKLRGFRVEPQEIEARLLAQDGVAQVAVLV 1554
             AG RLYRTGD AR+  DGVI+Y+GR+D QVK+RGFR+E  EIEARLL    V +  V+ 
Sbjct: 1987 VAGERLYRTGDLARYRPDGVIDYVGRIDHQVKVRGFRIELGEIEARLLEHAAVREAVVVA 2046

Query: 1555 RNTSAGPQLIGYYTAPEVAQGQD--ELNTRLKAALAAELPDYMVPAQLLRLDVMPLSPSG 1612
            +++ +G  L GY  A +  +  D   L   L A L   LP++MVP  L RL+ MPLSP+G
Sbjct: 2047 QDSRSGKVLSGYVVA-QAGEASDWPLLREALLAHLRLNLPEHMVPTHLTRLEHMPLSPNG 2105

Query: 1613 KLDRRALPEPQWQVREH--VEPASVLEQQIAGIWRQVLGLARIGLRDDFFALGGHSLLAT 1670
            KLDR+ALP P    R+     P + +E+ +  IW+ VLG+ R+G  D+FFALGG S+ + 
Sbjct: 2106 KLDRKALPAPDLAERQEAFTAPVNAIERALVDIWQGVLGVERVGTADNFFALGGDSINSI 2165

Query: 1671 QIISRTRQTCDVELPLRTLFEASELGAFAEQVRLIQASGQANRQPPIEKVDRSQPVPLSY 1730
            Q++SR R    + L  + LF    + A A +V      G  ++ P       S  VP+ +
Sbjct: 2166 QVVSRARAQ-GIGLTPKDLFLHQNIQALA-RVATAADEGVIDQGP---VTGDSLLVPMQH 2220

Query: 1731 SQQRMWFLWQMEPDSPAYNVGGMARLRGVLDVGRFEAALQALILRHETLRTTFPSV-DG- 1788
                 WF     P    +N   +   R  LD+   + ALQ ++  H+ LR  F    DG 
Sbjct: 2221 -----WFFASDIPHRDHWNQSVLLTPREPLDIAHLQQALQHVVAHHDALRLRFTQANDGR 2275

Query: 1789 --VAHQQVHGETGLRMDWKDFSMLAFDVREQRVQQLADSEAHKPFDLETGPLLRACLVKT 1846
               +HQ V      +   + +   A D  E  +  +AD +A +  DL+ GPLLR  L+  
Sbjct: 2276 WHASHQAVD-----KQPLEIWQRTAQDAAE--IIAIAD-QAQRSLDLQHGPLLRCSLIAV 2327

Query: 1847 AEHEHYFVLTLHHIVTEGWAMDIFARELSALYEAFIDDRESPLEPLPVQYLDYSVWQRQ- 1905
             +      L +HH+V +G +  +   +L  +Y      R +    LP +   +  W  Q 
Sbjct: 2328 QDGSSRLHLAIHHLVMDGVSWRVLLEDLQNVYRQL---RANQAPALPGKSTAFKAWAAQL 2384

Query: 1906 --WLESGERQRQLDYWTTQLGREHPLLELPGDRPRPSVQSHRGELFRFDLSDDLAARVRA 1963
              +  S    ++LDYW  QL  + PL  LP  RP  S  +  G      +S  L A    
Sbjct: 2385 QDYAHSETLNKELDYWVGQL--DQPLPPLPRARPEGSQAAAHGR----SVSTRLDATQTQ 2438

Query: 1964 FNAQQGLTLFMT-----MTAALAVLLYRYSGQTDLRIGAPVANRIRPESEGL-----IGA 2013
               +Q    + T     + +ALA ++ +++GQ    +      R     EG+     +G 
Sbjct: 2439 RLLKQAPAAYRTQVNDLLLSALAQVVCQWTGQPACLVQLEGHGR-EELFEGIDLSRTVGW 2497

Query: 2014 F--LNTQVLRCQLDGQMSVGELFEQVRHTVIEGQSHQDLPFDHLVEALQ------PPRSA 2065
            F  L   +L    D   S+  + EQ+R    +G  +  L +    EA+Q      PP   
Sbjct: 2498 FTSLFPVLLTPAGDPAESIKRIKEQLRAVPNKGLGYGVLRYLG-AEAVQQRLAKLPPARI 2556

Query: 2066 AYNPLFQVMCNVQRWEFQQSRTLAGMTVEYLVNDARATKFD------LNLEVTDLDQRLG 2119
             +N L Q         F QS     + V            D      L ++    + +L 
Sbjct: 2557 TFNYLGQ---------FDQSFDEQALFVPSEEGSGAGHGDDVALGNWLTIDGRIFNGQLT 2607

Query: 2120 CCLTYSTDLFDEPRIARMAGHWRNLLEALLD 2150
               T+S D+FDE  + R+A  +   L+ L+D
Sbjct: 2608 LDWTFSGDVFDEATLQRLADAYAVALQQLID 2638



 Score =  942 bits (2435), Expect = 0.0
 Identities = 600/1542 (38%), Positives = 843/1542 (54%), Gaps = 106/1542 (6%)

Query: 1727 PLSYSQQRMWFLWQMEPDSPAYNVGGMARLRGVLDVGRFEAALQALILRHETLRTTFPSV 1786
            PLS  Q  M F   +  DS    +  ++     LD  RF  A  ++I R + LRT+F   
Sbjct: 2678 PLSPMQDGMLF-HTLADDSSGLYINQISLPVHGLDTERFARAWTSVIEREDILRTSFHWQ 2736

Query: 1787 DGV-AHQQV---HGETGLRM-DWKDFSMLAFDVREQRVQQLADSEAHKPFDLETGPLLRA 1841
            DG+ A  Q+   H E  LR+ DW+       D+ EQ +   A ++    FDL   PL R 
Sbjct: 2737 DGLSAPVQIVHRHVELPLRIEDWR-----GQDISEQAISARATADYLAGFDLSRAPLQRV 2791

Query: 1842 CLVKTAEHEHYFVLTLHHIVTEGWAMD-IFARELSALYEAFIDDRESPLEPLPVQYLDYS 1900
             L++  +  +  + T HHI+ +GW+   +F   L       +  R         +Y ++ 
Sbjct: 2792 LLLRLEDDRYQMIWTCHHILMDGWSSSRLFGEVLQFYAHGHVPSRNG-------RYREFI 2844

Query: 1901 VW-QRQWLESGERQRQLDYWTTQLG---REHPLLELPGDRPRPSVQSHRGELFRFDLSDD 1956
             W QRQ  + G  +R   +W ++L    +   L +    R   ++  H     R+D++  
Sbjct: 2845 EWLQRQ--DQGALER---FWRSRLDTLEQATSLSQAMFPRHVEALPGHEALYSRWDVAQ- 2898

Query: 1957 LAARVRAFNAQQGLTLFMTMTAALAVLLYRYSGQTDLRIGAPVANRIRP--ESEGLIGAF 2014
              AR++       +T    +  A  +LL R++GQ  +  GA VA R      ++ ++G F
Sbjct: 2899 -TARLQQACRALQITPNTLIQGAWLLLLQRFTGQNTVVFGATVAGRPAGLVNADNILGLF 2957

Query: 2015 LNTQVLRCQLDGQMSVGELFEQVRHTVIEGQSHQDLPFDHLVEALQPPRSAAYNPLFQVM 2074
            +NT  +   LD +  + +  +Q++   ++ + H   P                      +
Sbjct: 2958 INTLPVIQTLDPEQPLDQWLQQLQAYNLDLREHSHAP----------------------L 2995

Query: 2075 CNVQRWE-------FQQSRTLAGMTVEYLVNDARATKFDL----NLEVT----DLDQRLG 2119
             ++QRW        F          ++  + D + T        N +VT    DL   LG
Sbjct: 2996 ADIQRWSGLGGQGLFDSIIVFENYPIDERLGDQQDTGLSFGESRNHDVTNFPMDLAVNLG 3055

Query: 2120 CCLT----YSTDLFDEPRIARMAGHWRNLLEALLDDPARRLSELPLLDTREQQQLLDSLG 2175
              L+    +  + F    + R+     + LEA++D P   L  L  L   EQ Q      
Sbjct: 3056 ETLSIEYLFLRNAFSVEAVERIRRTLEDTLEAMIDAPHVCLGNLQRLSL-EQWQAFAQWS 3114

Query: 2176 VEPGEHHLDRCLHELFDEQARMRPDAPALTFAGQTLTYAQLDSSANRLAWMLRERGVGPQ 2235
              P   +  + L EL    A+ RP A AL     +L Y +L+  ANRLA  L E G GP+
Sbjct: 3115 AAPNGCYQAQLLPELIGRHAQSRPQATALVCGDDSLDYGELERRANRLAHRLIELGAGPE 3174

Query: 2236 VRVGLALERSLEMVIGLLAILKAGGAYVPLDPEYPLDRLHYMIEDSGISLLLSDRAMLQA 2295
            V VG+ALERS++MV+ LLA++K+G AYVPLD +YP +RL +MIEDS ++LLL+   +   
Sbjct: 3175 VVVGVALERSVDMVVALLAVMKSGSAYVPLDIDYPPERLAFMIEDSAMALLLTHSRVQAR 3234

Query: 2296 LGALPDGVGCWCLED-DSPALAQYPDHALPLINLPQHQAYLIYTSGSTGKPKGVVVSHGE 2354
            L  L DG+    ++  DS   +  P H+  L     + AYLIYTSGSTG+PKGV V+HG 
Sbjct: 3235 LPDL-DGLARLAIDGFDSRGYSDLPPHSGLLGG---NLAYLIYTSGSTGRPKGVAVAHGP 3290

Query: 2355 IAMHCQAVIQRFGMRPDDCELHFYSINFDAATERLLVPLLSGAHVVLRAQGQWDAEEICG 2414
            ++MHCQA+   + M     ELHF S  FD A ER L  L SG  +V+R    W  E+   
Sbjct: 3291 MSMHCQAIAGLYDMDEQTRELHFMSFAFDGAHERWLSTLFSGGTLVIRDGALWTPEQTFD 3350

Query: 2415 LIRQHRINILGFTPSYGSQLAQWLATQGQTLP-VRMCITGGEALTGEHLQRIRAAFNPSL 2473
             + +H I I  F P+Y  QLA++ A      P VR+   GG+A+  +  ++++AA  P  
Sbjct: 3351 ALHRHGITIACFPPAYLKQLAEYAANSSLAPPLVRVYCFGGDAVPDQTFEQVKAALRPQF 3410

Query: 2474 FFNAYGPTETVVMPLASLAPEQLEEGAASVPIGSVIGARMAYILDADLALVPPGATGELY 2533
            F N YGPTETVV P+    P      AA  PIG  +G R  Y+LD +L  +PPG  GELY
Sbjct: 3411 FTNGYGPTETVVTPMLWKVPASERCEAAYAPIGRAVGERALYVLDDNLNPLPPGIAGELY 3470

Query: 2534 VGGAGLAQGYHQRPGMTAERFVADPFATDGGRLYRTGDLVRQRADGLVEYLGRIDHQVKI 2593
            +GGAG+A+GYH+RP ++AERFV DPF   G RLYR+GDLVRQR DG+++Y+GRIDHQ+K+
Sbjct: 3471 IGGAGIARGYHRRPDLSAERFVPDPFGQPGDRLYRSGDLVRQRVDGVLDYVGRIDHQIKV 3530

Query: 2594 RGFRIELGEIETRLLEHDAVREAVVLALDTPGGKQLAGYLVSDVASQDDGRQAALRDALK 2653
            RGFRIELGE+E  L + D V +A+V+A D+  GKQL GY+V+      DG   A+ D LK
Sbjct: 3531 RGFRIELGEVEASLRQLDDVSDALVIARDSASGKQLIGYVVTP-----DG--VAIGDRLK 3583

Query: 2654 NHLKAQLPDYMVPTHLILLASMPLTANGKLDRRALPMPDPELNRQHYVAPSNALEQTLAA 2713
              L+AQLPDYMVP  +I L ++P+T NGKLDR+ALP  +PE     YV   N  EQ LA 
Sbjct: 3584 AELRAQLPDYMVPAQVICLEALPVTPNGKLDRQALP--EPEFKGAEYVPGRNRDEQLLAE 3641

Query: 2714 IWCDVLNVTQVGLNDNFFELGGDSILSIQVVSRARQQG---IHFTPRDLFQHQTVQTLAA 2770
            +W +VL V +VG+ DNFFELGGDSILS+QVVSR R  G   +    RDL + QT++ +  
Sbjct: 3642 VWAEVLQVERVGITDNFFELGGDSILSLQVVSRVRNHGELKMELKLRDLMRGQTIEAIFD 3701

Query: 2771 VATRSQQVLAE---QGLLQGESGLTPIQHWFFERPIPNRHHWNQALVLEPTTALAPQPLE 2827
              + +  V  E   Q   +G+  L PIQ W+F++ +   HH+NQAL+L     L    LE
Sbjct: 3702 QRSNTFAVTVEDVTQVAAEGQFNLIPIQQWWFDQRMSEPHHYNQALMLRARQGLNLTALE 3761

Query: 2828 QALRSLLQHHDALRLGFDESTG----HWQA----EHQPLSDAALLRHVSVDTAQDCEALF 2879
             ALR ++Q HDALRL F E  G    H+QA    + Q  ++  L +H   D A+  E   
Sbjct: 3762 LALRGMMQQHDALRLRFSEVGGRHLQHYQALEEMQAQWANEPLLWQHSVADEAE-LELYA 3820

Query: 2880 AETQSRFNLATGPLLRAVLAQFPDGQQRLLIVIHHLVVDGVSWRVLMDDLQTVYRPLAIG 2939
             + Q   ++  GP+ R        G+ R+L+VIHHLV+D VSWR+L+ DLQT Y   + G
Sbjct: 3821 EQVQRSLDVHNGPVWRTAHVTLGSGEIRVLMVIHHLVIDTVSWRLLLQDLQTAYAAYSDG 3880

Query: 2940 QAAQLSAKTSPMRDWAARLQAYAGSESLREELAWWQAQLSGAQVPLPCANPEGGLLERHA 2999
            Q   L  +TS  R WA  LQAYA      E+ AWW  Q+      LPC N  G    ++ 
Sbjct: 3881 QTPNLPIRTSSYRAWAEGLQAYARDYPAPEK-AWWLEQIDQPGSELPCDNLRGKNQVQYQ 3939

Query: 3000 ETVSVRLDAERTRQLLQQAPSAYRTQVNDLLLTALARVLCRWSGHASTLVQLEGHGREAL 3059
                + L  + T+QLL+Q P+ Y+TQ+ND+LL AL+RVLCRWS   S L+QLEGHGRE L
Sbjct: 3940 SVAHLSLSEQHTQQLLKQVPAVYQTQINDVLLAALSRVLCRWSQQPSVLIQLEGHGREDL 3999

Query: 3060 FDDIDLTRTVGWFTSVYPVRLSPATEG-FGASIKAIKEQLRAVPHKGLGHGVLRYLADAA 3118
              DIDL+R++GWFTS++PVRL+P+ E   G S++A++ QL AVP+KGLG+GVLRY++   
Sbjct: 4000 VKDIDLSRSLGWFTSMFPVRLTPSHEADIGTSVRAVQRQLAAVPNKGLGYGVLRYMSGEP 4059

Query: 3119 TRQTMANLPHAAITFNYLGQLDQTLGHDALFQPLDAPLGAIHDPDAPLPNELSVDSQVSG 3178
               T+   P A +TFNYLGQ DQ+    AL  P     G+ + P A L N L V  QV  
Sbjct: 4060 MAATLGRAPQARVTFNYLGQFDQSFDDKALLVPAAESYGSCYSPHAALGNWLEVVGQVYD 4119

Query: 3179 GELLLRWTFSRERHDRQAIATLAGAYLDELQRLIEHCLTDEA 3220
            G L LR  FS  R+  + +  L  AY +EL+ LIEHC+   A
Sbjct: 4120 GRLALRCIFSTRRYRAETVEGLMRAYQEELEALIEHCVAQAA 4161



 Score =  609 bits (1571), Expect = e-177
 Identities = 451/1365 (33%), Positives = 678/1365 (49%), Gaps = 85/1365 (6%)

Query: 667  DEALPQSLAQNRLWITWRLDPTSHAYNIPGALHLRGELDEDALRTSFQQLIERHESLRTR 726
            ++  P S  Q+ +      D +S  Y    +L + G LD +    ++  +IER + LRT 
Sbjct: 2674 EDIYPLSPMQDGMLFHTLADDSSGLYINQISLPVHG-LDTERFARAWTSVIEREDILRTS 2732

Query: 727  FLERDGVA--LQQIDPAGEFKLHVIDLGDLPADSREARARQVREDEAATPFDLEKGPLLR 784
            F  +DG++  +Q +    E  L + D        +   AR   +  A   FDL + PL R
Sbjct: 2733 FHWQDGLSAPVQIVHRHVELPLRIEDWRGQDISEQAISARATADYLAG--FDLSRAPLQR 2790

Query: 785  VTLLRLDEDDHQLLVTLHHIIADGWSMSVLIEEFSRLYAAASQGQVAELAPLPLQYADYG 844
            V LLRL++D +Q++ T HHI+ DGWS S L  E  + YA    G V        +Y ++ 
Sbjct: 2791 VLLLRLEDDRYQMIWTCHHILMDGWSSSRLFGEVLQFYA---HGHVPSRNG---RYREFI 2844

Query: 845  SW-QRQWLAQGEAERQLSYWKQQLGA-EQSALVLGTDHPRSAQRQRSAARHSLRLDKDLG 902
             W QRQ   QG  ER   +W+ +L   EQ+  +     PR  +          R D    
Sbjct: 2845 EWLQRQ--DQGALER---FWRSRLDTLEQATSLSQAMFPRHVEALPGHEALYSRWDVAQT 2899

Query: 903  EALRHVAQAHDATPFMLLLAAFQALLQRYTGQSDIRIGVPNANRPR--LETQGLIGFFIN 960
              L+   +A   TP  L+  A+  LLQR+TGQ+ +  G   A RP   +    ++G FIN
Sbjct: 2900 ARLQQACRALQITPNTLIQGAWLLLLQRFTGQNTVVFGATVAGRPAGLVNADNILGLFIN 2959

Query: 961  TQVLRGQVDSRQPFAALLAQAREATLGAQAHQDLPFEQLLEAIPEAREQGLFQ---VMFN 1017
            T  +   +D  QP    L Q +   L  + H   P   + +       QGLF    V  N
Sbjct: 2960 TLPVIQTLDPEQPLDQWLQQLQAYNLDLREHSHAPLADI-QRWSGLGGQGLFDSIIVFEN 3018

Query: 1018 HQQRDLSALRRLPGLLAEELPWHS-REAKFDLQLHSEEDRNGRLTLSFDYADELFEAATI 1076
            +   +    ++  GL   E   H       DL ++  E     L++ + +    F    +
Sbjct: 3019 YPIDERLGDQQDTGLSFGESRNHDVTNFPMDLAVNLGET----LSIEYLFLRNAFSVEAV 3074

Query: 1077 ERLATHYGNLLRAVCAGPQQAIGDVPLLTADEHSQQQQWSAAP--CAPASQWLPELLNEQ 1134
            ER+     + L A+   P   +G++  L+ ++     QWSAAP  C  A Q LPEL+   
Sbjct: 3075 ERIRRTLEDTLEAMIDAPHVCLGNLQRLSLEQWQAFAQWSAAPNGCYQA-QLLPELIGRH 3133

Query: 1135 ARNTPQRIALRWDGGQLDYAELHAQANRLAHYLRDKGVGPDVCVAIAAERSPQLLIGVLA 1194
            A++ PQ  AL      LDY EL  +ANRLAH L + G GP+V V +A ERS  +++ +LA
Sbjct: 3134 AQSRPQATALVCGDDSLDYGELERRANRLAHRLIELGAGPEVVVGVALERSVDMVVALLA 3193

Query: 1195 IIKAGGAYVPLDADYPAERLAYMLDDSGVDLLLTQTALLERLPACDGVSVIAMDALHLEQ 1254
            ++K+G AYVPLD DYP ERLA+M++DS + LLLT + +  RLP  DG++ +A+D      
Sbjct: 3194 VMKSGSAYVPLDIDYPPERLAFMIEDSAMALLLTHSRVQARLPDLDGLARLAIDGFDSRG 3253

Query: 1255 WPSNPPGLHLHGDHLAYVIYTSGSTGQPKGVGNTHAALAERLQWMQATYGLDAIDVLMQK 1314
            +   PP   L G +LAY+IYTSGSTG+PKGV   H  ++   Q +   Y +D     +  
Sbjct: 3254 YSDLPPHSGLLGGNLAYLIYTSGSTGRPKGVAVAHGPMSMHCQAIAGLYDMDEQTRELHF 3313

Query: 1315 APISFDVSVWECFWPLITGSQLVLAGPGEHRDPHRIAQLVQQFGVTTLHFVPPLLSLFVD 1374
               +FD +       L +G  LV+   G    P +    + + G+T   F P  L    +
Sbjct: 3314 MSFAFDGAHERWLSTLFSGGTLVIR-DGALWTPEQTFDALHRHGITIACFPPAYLKQLAE 3372

Query: 1375 EPL-SARCTSLRRLFS-GGEALPAELRNRVLAQLPSVQLHNRYGPTETAINVTHWH---C 1429
                S+    L R++  GG+A+P +   +V A L      N YGPTET +    W     
Sbjct: 3373 YAANSSLAPPLVRVYCFGGDAVPDQTFEQVKAALRPQFFTNGYGPTETVVTPMLWKVPAS 3432

Query: 1430 TEADGERSPIGRPLGNVLCRVLDSDLNPVPAGVPGELCIGGLGLARGYLGRPGLSAERFV 1489
               +   +PIGR +G     VLD +LNP+P G+ GEL IGG G+ARGY  RP LSAERFV
Sbjct: 3433 ERCEAAYAPIGRAVGERALYVLDDNLNPLPPGIAGELYIGGAGIARGYHRRPDLSAERFV 3492

Query: 1490 ADPLGQAGARLYRTGDRARWNADGVIEYLGRLDQQVKLRGFRVEPQEIEARLLAQDGVAQ 1549
             DP GQ G RLYR+GD  R   DGV++Y+GR+D Q+K+RGFR+E  E+EA L   D V+ 
Sbjct: 3493 PDPFGQPGDRLYRSGDLVRQRVDGVLDYVGRIDHQIKVRGFRIELGEVEASLRQLDDVSD 3552

Query: 1550 VAVLVRNTSAGPQLIGYYTAPE-VAQGQDELNTRLKAALAAELPDYMVPAQLLRLDVMPL 1608
              V+ R++++G QLIGY   P+ VA G      RLKA L A+LPDYMVPAQ++ L+ +P+
Sbjct: 3553 ALVIARDSASGKQLIGYVVTPDGVAIGD-----RLKAELRAQLPDYMVPAQVICLEALPV 3607

Query: 1609 SPSGKLDRRALPEPQWQVREHVEPASVLEQQIAGIWRQVLGLARIGLRDDFFALGGHSLL 1668
            +P+GKLDR+ALPEP+++  E+V   +  EQ +A +W +VL + R+G+ D+FF LGG S+L
Sbjct: 3608 TPNGKLDRQALPEPEFKGAEYVPGRNRDEQLLAEVWAEVLQVERVGITDNFFELGGDSIL 3667

Query: 1669 ATQIISRTRQ--TCDVELPLRTLFEASELGAF-----------AEQVRLIQASGQANRQP 1715
            + Q++SR R      +EL LR L     + A             E V  + A GQ N  P
Sbjct: 3668 SLQVVSRVRNHGELKMELKLRDLMRGQTIEAIFDQRSNTFAVTVEDVTQVAAEGQFNLIP 3727

Query: 1716 PIEKVDRSQPVPLSYSQQRMWFLWQMEPDSPAYNVGGMARLRGVLDVGRFEAALQALILR 1775
                             Q+ WF  +M  +   YN   M R R  L++   E AL+ ++ +
Sbjct: 3728 ----------------IQQWWFDQRMS-EPHHYNQALMLRARQGLNLTALELALRGMMQQ 3770

Query: 1776 HETLRTTFPSVDGVAHQQVHGETGLRMDWKDFSML-AFDVREQRVQQLADSEAHKPFDLE 1834
            H+ LR  F  V G   Q       ++  W +  +L    V ++   +L   +  +  D+ 
Sbjct: 3771 HDALRLRFSEVGGRHLQHYQALEEMQAQWANEPLLWQHSVADEAELELYAEQVQRSLDVH 3830

Query: 1835 TGPLLRACLVKTAEHEHYFVLTLHHIVTEGWAMDIFARELSALYEAFIDDRESPLEPLPV 1894
             GP+ R   V     E   ++ +HH+V +  +  +  ++L   Y A+  D ++P   LP+
Sbjct: 3831 NGPVWRTAHVTLGSGEIRVLMVIHHLVIDTVSWRLLLQDLQTAYAAY-SDGQTP--NLPI 3887

Query: 1895 QYLDYSVWQRQWLESGERQ---RQLDYWTTQLGREHPLLELPGDRPRPSVQSHRGELFRF 1951
            +   Y  W  + L++  R     +  +W  Q+  + P  ELP D  R   Q     +   
Sbjct: 3888 RTSSYRAW-AEGLQAYARDYPAPEKAWWLEQI--DQPGSELPCDNLRGKNQVQYQSVAHL 3944

Query: 1952 DLSDDLAAR-VRAFNAQQGLTLFMTMTAALAVLLYRYSGQTDLRI 1995
             LS+    + ++   A     +   + AAL+ +L R+S Q  + I
Sbjct: 3945 SLSEQHTQQLLKQVPAVYQTQINDVLLAALSRVLCRWSQQPSVLI 3989



 Score =  497 bits (1280), Expect = e-143
 Identities = 370/1124 (32%), Positives = 554/1124 (49%), Gaps = 87/1124 (7%)

Query: 3240 SLPVPAGQIE-DVYPLTPMQEGMLLHTLLEPGTGLYYMQDRYRINSELDPQRFAQAWQAV 3298
            +LP+P    E D   L+  QE       L+P +  Y+M    R+   LD     Q++ A+
Sbjct: 37   NLPIPPVSAEFDTLSLSFAQERQWFLWQLDPESAAYHMPTALRLRGPLDIVALEQSFTAL 96

Query: 3299 IARHEALRASFCWNVGEDMLQIIHRPGSTPVDYLDWSEVPEDAQEAKLQALLKNEREAGF 3358
            +ARHE+LR +F  + GE ++Q +H      +      ++P++ Q   L+ L++ E    F
Sbjct: 97   VARHESLRTTFQTH-GEQVVQQVHAAAPVRLQADTLEQLPDEMQ---LRQLIEGETRQLF 152

Query: 3359 DLLNQAPFHLRLIRVGAARYWFMMSNHHILIDAWCRSLLMDDFFEIYTALGENREPQL-T 3417
            DL        RL+R+ A  +  +++ HHI+ D W   +++++  + Y A  + R PQL  
Sbjct: 153  DLQQGPLLRTRLLRLAADDHVLIVTLHHIVADGWSMQVMVEELVDGYAAFSQGRTPQLPA 212

Query: 3418 VPPRYRDYIGWLQHQSLAEARQ----WWQRNLQGFERTTPIPGD--RPFLREHAGDSGGM 3471
            +P +Y DY  W +H   A  R+    +W   L G +    +P D  RP +  + G    +
Sbjct: 213  LPIQYTDYAIWQRHWMEAGERERQLEYWTARLGGQQPMLELPTDFPRPAVMSYRGARLEL 272

Query: 3472 IVGDCYTRLDARDGARLRELAQAHQLTVNTFAQAAWALVLRRVSGDRDVLFGVTVAGRPV 3531
             +       DA     L+ +AQ   +++     A++  +L R SG  D+  GV  A R  
Sbjct: 273  AI-------DAALAGDLKHMAQREGVSLFMLLLASFQALLHRYSGQNDIRVGVPTANR-- 323

Query: 3532 ELPQMQRTVGLFINSIALRVQMPEDHQRCSVRQWLSDLLDHNMQLREYEYLPLVSIQEAS 3591
               + +R +G F+N+  L   +       S    L  +    +  + ++ LP   + EA 
Sbjct: 324  NRVEAERLIGFFVNTQVLDSDVQSQD---SFGALLQQVKQTALGAQAHQELPFEQLVEAL 380

Query: 3592 ELPKG---QPLFDSLFVFEN-APVEVSVLDRAQSLNATSDSGRTHTNFPLTAVCYPGDDL 3647
            +  +     PLF  +F      P     +   +      DS     +  L  V +  + L
Sbjct: 381  QPERNLSHSPLFQVMFNHRTQGPAGEGPVGALKVEELRWDSHTAQFDLTLETVEH-SEGL 439

Query: 3648 GLHLSYDQRYFDEATVQGLLSEFKRLLLALMEGLHGDMSELPLIGAQEQDFLIEGCNQSE 3707
               L Y    F   T+  +   ++ LL A++      + ELPL+ A E+   ++   ++ 
Sbjct: 440  SAGLIYATDLFGAETIARMAGHWQNLLRAIVADPAQRIGELPLLDAAERQAQLDNGQRTP 499

Query: 3708 HAYPLEQSYVELFEARVAAHPHRPAVSCLDTSYSYAELNARSNRLGHALIAAGVGLDQPV 3767
               P      +LFE + A  P  PA+      +SY +LNAR+N+L   L + GVG D  V
Sbjct: 500  TPAPEAGFVHQLFERQAARTPDAPALVIDGERWSYGQLNARANQLAGKLQSLGVGPDVLV 559

Query: 3768 ALLAERDAELLGMIVGSFKAGAGYLPLDPGLPSQRLGRIIELSRTPLLVCTQACQAQAQA 3827
             +  ER A +L  ++   KAG  YLPLDP  P  RL  +IE S   LL+ TQ+      +
Sbjct: 560  GIAVERSAHMLVGLLAILKAGGAYLPLDPAFPQDRLAYMIEDSGIELLL-TQS------S 612

Query: 3828 LLDEFACSGRPKL-----------LVWEAVQASALS------------------------ 3852
            LLD+   S   +L           +  E + AS L+                        
Sbjct: 613  LLDQLRLSCGSELARDSGGSVNIDVGGEGLIASKLAPTVFGVGVTTLVLDQEGDWLDGYA 672

Query: 3853 LENPGVYSGPDNLAYVIYTSGSTGLPKGVMVEQRGMLNNQLSKVPYLALSEVDVIAQTAS 3912
             E P      ++LAY IYTSGSTG PKGVMV    + N   S      ++  D      +
Sbjct: 673  CEAPAPRLDAEHLAYAIYTSGSTGKPKGVMVRHGALTNFVASMAQAPGMTAQDRALSLTT 732

Query: 3913 QSFDISVWQFLAAPLFGARVDIVPNAIAHDPQGLLEHVQQQGITVLESVPSLIQGMLAQE 3972
             SFDI   +     + GA + +    +A DP  +L  +  + + V+++ PS  + +L  E
Sbjct: 733  FSFDIFGLEIYVPLMVGACIVLTGQHVAQDPYAVLALIAAEQVNVVQATPSTWRMLLDNE 792

Query: 3973 RI-GLDGLRWMLPTGEAMPPELAHQWLLRYPDIGLVNAYGPAECSDDVAFFRVDLASTRG 4031
               GL G +  L  GEA+P ELA + L    ++   N YGP E +   A   +    T+ 
Sbjct: 793  HAQGLRGCK-CLCGGEALPQELAVRMLALGGEVW--NLYGPTETTIWSALHPLQ---TQA 846

Query: 4032 TYLPIGTPTDNNRLYLLDGALDLVPLGAVGELCVAGIGVGRGYVSDPLRTAPVFVPNPFG 4091
                +G P DN  LY++   L  VP G  GEL + G G+ RGY   P  TA  FVP+PF 
Sbjct: 847  AQPWLGGPIDNTALYIVGADLMPVPYGVAGELLIGGQGLARGYFQRPALTAERFVPHPFV 906

Query: 4092 APGERLYRTGDLARRRSDGVLEYVGRIDHQVKIRGYRIELGEIEARLHEQPEVRDGAVGV 4151
            A GERLYRTGDLAR+R+DG+LEY+GRIDHQVKIRG+RIELGEIEARL EQ  VR+ AV  
Sbjct: 907  A-GERLYRTGDLARQRADGILEYLGRIDHQVKIRGFRIELGEIEARLLEQDSVREAAVLA 965

Query: 4152 QEGVNGKHLVGYLVAT------DSALKPAERLERIKQRLRAELPEYMVPLHWLWLERLPL 4205
            QE   G+ LV Y+V T      D+  + A R + +  RL+  LP+YMVP H + L R PL
Sbjct: 966  QEAAGGQALVAYIVPTAELNDLDAGAQGALR-DSLTARLKVNLPDYMVPAHMVVLPRFPL 1024

Query: 4206 NANGKLDRKALPALEIGQLQSQDYQAPGNELEQTLADIWAQVLKVERVGVRDNFFELGGH 4265
              NGKLDRK LP  +  Q Q + Y AP   ++Q +A IWA +LKV +VG+ D+FFELGGH
Sbjct: 1025 TPNGKLDRKQLPKPDASQWQ-KTYVAPQTPVQQAVATIWADILKVPQVGLDDSFFELGGH 1083

Query: 4266 SLLATQIASRVQKALQRNVPLRAMFECSTVQELADYIDGLAASE 4309
            SLLATQ+ SR+++ L   VPLR +F  ++++   + +   +A +
Sbjct: 1084 SLLATQVMSRIRQQLNIQVPLRLLFAHNSLRAFVEAMGTASAHD 1127



 Score =  197 bits (500), Expect = 8e-53
 Identities = 244/935 (26%), Positives = 393/935 (42%), Gaps = 111/935 (11%)

Query: 41   LSYRDLDLRARTIAAALQAEAALGDRAVL--LFPSGPDYVAAFFGCLYAGVIAVPA---Y 95
            L Y +L+ RA  +A  L  E   G   V+        D V A    + +G   VP    Y
Sbjct: 3150 LDYGELERRANRLAHRL-IELGAGPEVVVGVALERSVDMVVALLAVMKSGSAYVPLDIDY 3208

Query: 96   PPESTRRHHQERLLSIIADAEPRLLLTRSDLREPLQAIESAPPVLCVDTLDPSRAGQWRM 155
            PPE        RL  +I D+   LLLT S ++  L  ++     L +D  D    G   +
Sbjct: 3209 PPE--------RLAFMIEDSAMALLLTHSRVQARLPDLDGLAR-LAIDGFDSR--GYSDL 3257

Query: 156  PT---LQDDDIAFLQYTSGSTALPKGVQVSHGNLVANELLIRHGFGIDLNPDDVIVSWLP 212
            P    L   ++A+L YTSGST  PKGV V+HG +  +   I   + +D    ++   ++ 
Sbjct: 3258 PPHSGLLGGNLAYLIYTSGSTGRPKGVAVAHGPMSMHCQAIAGLYDMDEQTREL--HFMS 3315

Query: 213  LYHDMGLIGGLLQPVFSGVPCVLMSPAYFLARPLRWLEAISEYGGTISGGPDFAYRLCSE 272
               D G     L  +FSG   V+   A  L  P +  +A+  +G TI+  P    +  +E
Sbjct: 3316 FAFD-GAHERWLSTLFSGGTLVIRDGA--LWTPEQTFDALHRHGITIACFPPAYLKQLAE 3372

Query: 273  RVSDSALQRLDLSGWRVAYSGSEPIRLDTLERFAEKFAPCGFTPDNFMASYGLAEATLFV 332
              ++S+L    +   RV   G + +   T E+      P  FT       YG  E  +  
Sbjct: 3373 YAANSSLAPPLV---RVYCFGGDAVPDQTFEQVKAALRPQFFTN-----GYGPTETVV-- 3422

Query: 333  AGTPRRHGIPSLRVDDAALAQNRAEPGQGSAVMSCGVSQPGHAVLIVEPATAQPLADNQV 392
              TP    +P+    +AA A        G AV        G   L V      PL     
Sbjct: 3423 --TPMLWKVPASERCEAAYAPI------GRAV--------GERALYVLDDNLNPLPPGIA 3466

Query: 393  GEVWAAGPSIALGYWRNPEASAKTFVQ----HDGRTWLRTGDLGFLR-DGELFITGRLKD 447
            GE++  G  IA GY R P+ SA+ FV       G    R+GDL   R DG L   GR+  
Sbjct: 3467 GELYIGGAGIARGYHRRPDLSAERFVPDPFGQPGDRLYRSGDLVRQRVDGVLDYVGRIDH 3526

Query: 448  LLIVRGHNLYPQDIEQAIEREVEVVRKGRVAAFAVNDGGE--------EGIGIAAEISRS 499
             + VRG  +   ++E ++ R+++ V    V A     G +        +G+ I   +   
Sbjct: 3527 QIKVRGFRIELGEVEASL-RQLDDVSDALVIARDSASGKQLIGYVVTPDGVAIGDRLKAE 3585

Query: 500  VQKILPPEALIKAIRQAVAEACQQAPSVVALLNPGALPKTSSGKLQRSACRSRLADGSLD 559
            ++  LP                   P+ V  L   ALP T +GKL R A           
Sbjct: 3586 LRAQLPD---------------YMVPAQVICLE--ALPVTPNGKLDRQA----------- 3617

Query: 560  SYAQFPSAEVEDADESAPSSELQTLIGQIWREQLQVEQVQADDHFFLLGGNSIAATQMIA 619
                 P  E + A+     +  + L+ ++W E LQVE+V   D+FF LGG+SI + Q+++
Sbjct: 3618 ----LPEPEFKGAEYVPGRNRDEQLLAEVWAEVLQVERVGITDNFFELGGDSILSLQVVS 3673

Query: 620  RLRE--ILGLELNLRLLFEAPTLAAFAEAVANQQQDGGQPQGAISALSRDEALPQSLAQN 677
            R+R    L +EL LR L    T+ A  +  +N      +    ++A  +   +P      
Sbjct: 3674 RVRNHGELKMELKLRDLMRGQTIEAIFDQRSNTFAVTVEDVTQVAAEGQFNLIP----IQ 3729

Query: 678  RLWITWRLDPTSHAYNIPGALHLRGELDEDALRTSFQQLIERHESLRTRFLERDGVALQQ 737
            + W   R+    H YN    L  R  L+  AL  + + ++++H++LR RF E  G  LQ 
Sbjct: 3730 QWWFDQRMSEPHH-YNQALMLRARQGLNLTALELALRGMMQQHDALRLRFSEVGGRHLQH 3788

Query: 738  IDPAGEFKLHVIDLGDLPADSREARAR-QVREDEAATPFDLEKGPLLRVTLLRLDEDDHQ 796
                 E +    +   L   S    A  ++  ++     D+  GP+ R   + L   + +
Sbjct: 3789 YQALEEMQAQWANEPLLWQHSVADEAELELYAEQVQRSLDVHNGPVWRTAHVTLGSGEIR 3848

Query: 797  LLVTLHHIIADGWSMSVLIEEFSRLYAAASQGQVAELAPLPLQYADYGSWQR--QWLAQG 854
            +L+ +HH++ D  S  +L+++    YAA S GQ      LP++ + Y +W    Q  A+ 
Sbjct: 3849 VLMVIHHLVIDTVSWRLLLQDLQTAYAAYSDGQTPN---LPIRTSSYRAWAEGLQAYARD 3905

Query: 855  EAERQLSYWKQQLGAEQSALVLGTDHPRSAQRQRSAARHSLRLDKDLGEALRHVAQAHDA 914
                + ++W +Q+    S L       ++  + +S A  SL  ++   + L+ V   +  
Sbjct: 3906 YPAPEKAWWLEQIDQPGSELPCDNLRGKNQVQYQSVAHLSLS-EQHTQQLLKQVPAVYQT 3964

Query: 915  TPFMLLLAAFQALLQRYTGQSDIRIGVPNANRPRL 949
                +LLAA   +L R++ Q  + I +    R  L
Sbjct: 3965 QINDVLLAALSRVLCRWSQQPSVLIQLEGHGREDL 3999



 Score =  172 bits (436), Expect = 2e-45
 Identities = 281/1133 (24%), Positives = 445/1133 (39%), Gaps = 155/1133 (13%)

Query: 13   HALQRRAALTPDRVALRFLADTPGQGVVLSYRDLDLRARTIAAALQAEAALGDRAV-LLF 71
            H ++ +    P   AL F      +G  LSY  L+ RA  +A AL       D  V +  
Sbjct: 1600 HLIEAQVERDPHAEALVF------EGQSLSYAQLNARANQLAHALIERGVGPDVLVGISV 1653

Query: 72   PSGPDYVAAFFGCLYAGVIAVPA---YPPESTRRHHQERLLSIIADAEPRLLLT-RSDLR 127
                + V      L AG   VP    YPPE        RL  +I D+   LLLT  S+L 
Sbjct: 1654 QRSVEMVVGLLAILKAGGAYVPLDPEYPPE--------RLAYMIEDSGIELLLTCGSELA 1705

Query: 128  EPLQA---IESAPPVLCVDTLDPSRAGQWRMPTLQDDD------------------IAFL 166
                    I+     L    L P+  G   +    DDD                  +A++
Sbjct: 1706 RDSGGSVNIDVEGEGLIASKLAPTGGGLNLLVLDGDDDWLSDHSTDNPPARAVAQNLAYV 1765

Query: 167  QYTSGSTALPKGVQVSHGNLVANELLIRHGFGIDLNPDDVIVSWLPLYHDMGLIGGLLQP 226
             YTSGST  PKG   SHG L      ++  +G++    D ++   P   D+  +     P
Sbjct: 1766 IYTSGSTGKPKGAGNSHGALTNRLCWMQQAYGLEAG--DSVLQKTPFSFDVS-VWEFFWP 1822

Query: 227  VFSGVPCVLMSPAYFLARPLRWLEAISEYGGTISGGPDFAYRLCSERVSDSALQRLDLSG 286
            + +GV  V+ +P      P + +  I     T      F   +    V D+ + R     
Sbjct: 1823 LMTGVRLVVAAPGDH-RDPQKLVSLIERESITTL---HFVPSMLQAFVLDANVSRCTCL- 1877

Query: 287  WRVAYSGSEPIRLDTLERFAEKFAPCGFTPDNFMASYGLAEATLFVAGTPRRHGIPSLRV 346
             R+  SG E + +D  ++   K              YG  EA + V     R        
Sbjct: 1878 TRIVCSG-EALPVDAQQQVFAKLPNA-----RLYNLYGPTEAAIDVTHWTCR-------- 1923

Query: 347  DDAALAQNRAEPGQGSAVMSCGVSQPGHAVLIVEPATAQPLADNQVGEVWAAGPSIALGY 406
            D+   A    EP               + +  +      P+     GE++ +G  +A GY
Sbjct: 1924 DEGRDAVPIGEP-------------IANLMTYILDDELNPVPSGVNGELYLSGQGLARGY 1970

Query: 407  WRNPEASAKTFVQHD---GRTWLRTGDLGFLR-DGELFITGRLKDLLIVRGHNLYPQDIE 462
             R PE +A+ FV      G    RTGDL   R DG +   GR+   + VRG  +   +IE
Sbjct: 1971 HRRPELTAERFVASPFVAGERLYRTGDLARYRPDGVIDYVGRIDHQVKVRGFRIELGEIE 2030

Query: 463  QAIEREVEVVRKGRVAAFAVNDGGEEGIGIAAEISRSVQKILPPEALIKAIRQAVAEACQ 522
              +  E   VR+  V A     G      + A+   +    L  EAL+  +R  + E   
Sbjct: 2031 ARL-LEHAAVREAVVVAQDSRSGKVLSGYVVAQAGEASDWPLLREALLAHLRLNLPE--H 2087

Query: 523  QAPSVVALLNPGALPKTSSGKLQRSACRSRLADGSLDSYAQFPSAEVEDADE--SAPSSE 580
              P+ +  L    +P + +GKL R A                P+ ++ +  E  +AP + 
Sbjct: 2088 MVPTHLTRLE--HMPLSPNGKLDRKA---------------LPAPDLAERQEAFTAPVNA 2130

Query: 581  LQTLIGQIWREQLQVEQVQADDHFFLLGGNSIAATQMIARLREILGLELNLRLLFEAPTL 640
            ++  +  IW+  L VE+V   D+FF LGG+SI + Q+++R R   G+ L  + LF    +
Sbjct: 2131 IERALVDIWQGVLGVERVGTADNFFALGGDSINSIQVVSRAR-AQGIGLTPKDLFLHQNI 2189

Query: 641  AAFAEAVANQQQDGGQPQGAISALSRDEALPQSLAQNRLWITWRLDPTSHAYNIPGALHL 700
             A A  VA    +G   QG ++  S        L   + W      P    +N    L  
Sbjct: 2190 QALAR-VATAADEGVIDQGPVTGDS-------LLVPMQHWFFASDIPHRDHWNQSVLLTP 2241

Query: 701  RGELDEDALRTSFQQLIERHESLRTRFLERDGVALQQIDPAGEFKLHVIDLGDLPADSRE 760
            R  LD   L+ + Q ++  H++LR RF + +       D         +D   L    R 
Sbjct: 2242 REPLDIAHLQQALQHVVAHHDALRLRFTQAN-------DGRWHASHQAVDKQPLEIWQRT 2294

Query: 761  AR-ARQV--REDEAATPFDLEKGPLLRVTLLRLDEDDHQLLVTLHHIIADGWSMSVLIEE 817
            A+ A ++    D+A    DL+ GPLLR +L+ + +   +L + +HH++ DG S  VL+E+
Sbjct: 2295 AQDAAEIIAIADQAQRSLDLQHGPLLRCSLIAVQDGSSRLHLAIHHLVMDGVSWRVLLED 2354

Query: 818  FSRLYAAASQGQVAELAPLPLQYADYGSWQRQWLAQGEAERQLSYWKQQLGAEQSALVLG 877
               +Y      Q   L      +  + +  + +       ++L YW  QL  +Q    L 
Sbjct: 2355 LQNVYRQLRANQAPALPGKSTAFKAWAAQLQDYAHSETLNKELDYWVGQL--DQPLPPLP 2412

Query: 878  TDHPRSAQRQRSAARHSLRLDKDLGEALRHVAQAHDATPFM-LLLAAFQALLQRYTGQSD 936
               P  +Q        S RLD    + L   A A   T    LLL+A   ++ ++TGQ  
Sbjct: 2413 RARPEGSQAAAHGRSVSTRLDATQTQRLLKQAPAAYRTQVNDLLLSALAQVVCQWTGQPA 2472

Query: 937  IRIGVPNANRPRL----ETQGLIGFFIN--TQVLRGQVDSRQPFAALLAQAREAT----- 985
              + +    R  L    +    +G+F +    +L    D  +    +  Q R        
Sbjct: 2473 CLVQLEGHGREELFEGIDLSRTVGWFTSLFPVLLTPAGDPAESIKRIKEQLRAVPNKGLG 2532

Query: 986  ------LGAQAHQDLPFEQLLEAIPEAREQGLFQVMFNH-QQRDLSALRRLPGLLAEE-- 1036
                  LGA+A Q     Q L  +P AR      + FN+  Q D S   +   + +EE  
Sbjct: 2533 YGVLRYLGAEAVQ-----QRLAKLPPAR------ITFNYLGQFDQSFDEQALFVPSEEGS 2581

Query: 1037 LPWHSREAKFDLQLHSE-EDRNGRLTLSFDYADELFEAATIERLATHYGNLLR 1088
               H  +      L  +    NG+LTL + ++ ++F+ AT++RLA  Y   L+
Sbjct: 2582 GAGHGDDVALGNWLTIDGRIFNGQLTLDWTFSGDVFDEATLQRLADAYAVALQ 2634



 Score = 48.1 bits (113), Expect = 6e-08
 Identities = 99/456 (21%), Positives = 170/456 (37%), Gaps = 46/456 (10%)

Query: 3249 EDVYPLTPMQEGMLLHTLLEPGTGLYYMQDRYRINSELDPQRFAQAWQAVIARHEALRAS 3308
            E  + L P+Q+      + EP    Y      R    L+      A + ++ +H+ALR  
Sbjct: 3720 EGQFNLIPIQQWWFDQRMSEPHH--YNQALMLRARQGLNLTALELALRGMMQQHDALRLR 3777

Query: 3309 FCWNVGEDMLQIIHRPGSTPVDYLDWSEVP-----EDAQEAKLQALLKNEREAGFDLLNQ 3363
            F   VG   LQ  H      +    W+  P       A EA+L+ L   + +   D+ N 
Sbjct: 3778 FS-EVGGRHLQ--HYQALEEMQ-AQWANEPLLWQHSVADEAELE-LYAEQVQRSLDVHNG 3832

Query: 3364 APFHLRLIRVGAARYWFMMSNHHILIDAWCRSLLMDDFFEIYTALGENREPQLTVPPRYR 3423
              +    + +G+     +M  HH++ID     LL+ D    Y A  + + P L  P R  
Sbjct: 3833 PVWRTAHVTLGSGEIRVLMVIHHLVIDTVSWRLLLQDLQTAYAAYSDGQTPNL--PIRTS 3890

Query: 3424 DYIGWLQ------HQSLAEARQWWQRNLQGFERTTPIPGDRPFLREHAGDSGGMIVGDCY 3477
             Y  W +          A  + WW   +       P    R   +        + + + +
Sbjct: 3891 SYRAWAEGLQAYARDYPAPEKAWWLEQIDQPGSELPCDNLRGKNQVQYQSVAHLSLSEQH 3950

Query: 3478 TRLDARDGARLRELAQAHQLTVNTFAQAAWALVLRRVSGDRDVLFGVTVAGRP--VELPQ 3535
            T+        L+++   +Q  +N    AA + VL R S    VL  +   GR   V+   
Sbjct: 3951 TQ------QLLKQVPAVYQTQINDVLLAALSRVLCRWSQQPSVLIQLEGHGREDLVKDID 4004

Query: 3536 MQRTVGLFINSIALRVQMPEDHQ-RCSVRQWLSDLLDHNMQLREYEYLPLVSIQEAS--- 3591
            + R++G F +   +R+    +     SVR     L     +   Y  L  +S +  +   
Sbjct: 4005 LSRSLGWFTSMFPVRLTPSHEADIGTSVRAVQRQLAAVPNKGLGYGVLRYMSGEPMAATL 4064

Query: 3592 -ELPKGQPLFDSLFVFENAPVEVSVLDRAQSLNATSDSGRTHTN-------FPLTAVCYP 3643
               P+ +  F+ L  F+      S  D+A  + A    G  ++          +    Y 
Sbjct: 4065 GRAPQARVTFNYLGQFDQ-----SFDDKALLVPAAESYGSCYSPHAALGNWLEVVGQVYD 4119

Query: 3644 GDDLGLHLSYDQRYFDEATVQGLLSEFKRLLLALME 3679
            G  L L   +  R +   TV+GL+  ++  L AL+E
Sbjct: 4120 G-RLALRCIFSTRRYRAETVEGLMRAYQEELEALIE 4154