Experiment: Survival; Treatment=UGA synthetic gw pH3.9 with 1 mM Glucose and 5 mM NaH2PO4; TreatmentDuration=24hrs; Outgrowth=LBkan50
Only strains with sufficient reads to estimate fitness are shown,
but the strain fitness values are still rather noisy and may be biased towards zero.
Strains near the edge of a gene are not shown as being associated with that
gene (they are in grey).
Strains in the central 10-90% of a gene are color coded by the insertion's strand.
Usually, "+" means that the selectable marker is encoded on the forward strand,
i.e., transcribed rightward.
500 nt IAI47_04630 and IAI47_04635 overlap by 4 nucleotides IAI47_04635 and IAI47_04640 overlap by 1 nucleotides IAI47_04640 and IAI47_04645 are separated by 15 nucleotides
IAI47_04630: IAI47_04630 - allose kinase, at 975,226 to 976,122
_04630
IAI47_04635: IAI47_04635 - ribulose-phosphate 3-epimerase, at 976,119 to 976,811
_04635
IAI47_04640: IAI47_04640 - ABC transporter permease, at 976,811 to 977,860
_04640
IAI47_04645: IAI47_04645 - ABC transporter permease, at 977,876 to 978,865
_04645
IAI47_04650: IAI47_04650 - sugar ABC transporter ATP-binding protein, at 978,858 to 980,363
_04650
Position (kb) 976
977
978 Strain fitness (log2 ratio) -3
-2
-1
0
1
2
3
4 at 975.873 kb on + strand, within IAI47_04630 at 975.874 kb on - strand, within IAI47_04630 at 975.933 kb on + strand, within IAI47_04630 at 975.933 kb on + strand, within IAI47_04630 at 975.934 kb on - strand, within IAI47_04630 at 975.934 kb on - strand, within IAI47_04630 at 975.934 kb on - strand, within IAI47_04630 at 975.934 kb on - strand, within IAI47_04630 at 975.972 kb on + strand, within IAI47_04630 at 975.972 kb on + strand, within IAI47_04630 at 975.972 kb on + strand, within IAI47_04630 at 975.972 kb on + strand, within IAI47_04630 at 975.973 kb on - strand, within IAI47_04630 at 975.973 kb on - strand, within IAI47_04630 at 975.973 kb on - strand, within IAI47_04630 at 976.188 kb on + strand at 976.205 kb on + strand, within IAI47_04635 at 976.205 kb on + strand, within IAI47_04635 at 976.206 kb on - strand, within IAI47_04635 at 976.206 kb on - strand, within IAI47_04635 at 976.210 kb on - strand, within IAI47_04635 at 976.210 kb on - strand, within IAI47_04635 at 976.210 kb on - strand, within IAI47_04635 at 976.210 kb on - strand, within IAI47_04635 at 976.253 kb on + strand, within IAI47_04635 at 976.253 kb on + strand, within IAI47_04635 at 976.253 kb on + strand, within IAI47_04635 at 976.253 kb on + strand, within IAI47_04635 at 976.253 kb on + strand, within IAI47_04635 at 976.253 kb on + strand, within IAI47_04635 at 976.254 kb on - strand, within IAI47_04635 at 976.264 kb on + strand, within IAI47_04635 at 976.264 kb on + strand, within IAI47_04635 at 976.264 kb on + strand, within IAI47_04635 at 976.265 kb on - strand, within IAI47_04635 at 976.265 kb on - strand, within IAI47_04635 at 976.277 kb on + strand, within IAI47_04635 at 976.294 kb on + strand, within IAI47_04635 at 976.303 kb on + strand, within IAI47_04635 at 976.303 kb on + strand, within IAI47_04635 at 976.303 kb on + strand, within IAI47_04635 at 976.303 kb on + strand, within IAI47_04635 at 976.303 kb on + strand, within IAI47_04635 at 976.303 kb on + strand, within IAI47_04635 at 976.303 kb on + strand, within IAI47_04635 at 976.304 kb on - strand, within IAI47_04635 at 976.304 kb on - strand, within IAI47_04635 at 976.304 kb on - strand, within IAI47_04635 at 976.304 kb on - strand, within IAI47_04635 at 976.304 kb on - strand, within IAI47_04635 at 976.304 kb on - strand, within IAI47_04635 at 976.304 kb on - strand, within IAI47_04635 at 976.304 kb on - strand, within IAI47_04635 at 976.309 kb on + strand, within IAI47_04635 at 976.368 kb on - strand, within IAI47_04635 at 976.429 kb on - strand, within IAI47_04635 at 976.430 kb on + strand, within IAI47_04635 at 976.430 kb on + strand, within IAI47_04635 at 976.431 kb on - strand, within IAI47_04635 at 976.431 kb on - strand, within IAI47_04635 at 976.431 kb on - strand, within IAI47_04635 at 976.431 kb on - strand, within IAI47_04635 at 976.432 kb on + strand, within IAI47_04635 at 976.432 kb on + strand, within IAI47_04635 at 976.432 kb on + strand, within IAI47_04635 at 976.432 kb on + strand, within IAI47_04635 at 976.432 kb on + strand, within IAI47_04635 at 976.432 kb on + strand, within IAI47_04635 at 976.432 kb on + strand, within IAI47_04635 at 976.432 kb on + strand, within IAI47_04635 at 976.433 kb on - strand, within IAI47_04635 at 976.433 kb on - strand, within IAI47_04635 at 976.433 kb on - strand, within IAI47_04635 at 976.433 kb on - strand, within IAI47_04635 at 976.433 kb on - strand, within IAI47_04635 at 976.433 kb on - strand, within IAI47_04635 at 976.433 kb on - strand, within IAI47_04635 at 976.433 kb on - strand, within IAI47_04635 at 976.433 kb on - strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.442 kb on + strand, within IAI47_04635 at 976.443 kb on - strand, within IAI47_04635 at 976.443 kb on - strand, within IAI47_04635 at 976.443 kb on - strand, within IAI47_04635 at 976.443 kb on - strand, within IAI47_04635 at 976.444 kb on + strand, within IAI47_04635 at 976.444 kb on + strand, within IAI47_04635 at 976.444 kb on + strand, within IAI47_04635 at 976.444 kb on + strand, within IAI47_04635 at 976.444 kb on + strand, within IAI47_04635 at 976.444 kb on + strand, within IAI47_04635 at 976.444 kb on + strand, within IAI47_04635 at 976.445 kb on - strand, within IAI47_04635 at 976.445 kb on - strand, within IAI47_04635 at 976.445 kb on - strand, within IAI47_04635 at 976.445 kb on - strand, within IAI47_04635 at 976.445 kb on - strand, within IAI47_04635 at 976.492 kb on + strand, within IAI47_04635 at 976.493 kb on - strand, within IAI47_04635 at 976.493 kb on - strand, within IAI47_04635 at 976.493 kb on - strand, within IAI47_04635 at 976.508 kb on + strand, within IAI47_04635 at 976.508 kb on + strand, within IAI47_04635 at 976.509 kb on - strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.556 kb on + strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.557 kb on - strand, within IAI47_04635 at 976.558 kb on + strand, within IAI47_04635 at 976.558 kb on + strand, within IAI47_04635 at 976.558 kb on + strand, within IAI47_04635 at 976.559 kb on - strand, within IAI47_04635 at 976.559 kb on - strand, within IAI47_04635 at 976.559 kb on - strand, within IAI47_04635 at 976.559 kb on - strand, within IAI47_04635 at 976.559 kb on - strand, within IAI47_04635 at 976.559 kb on - strand, within IAI47_04635 at 976.559 kb on - strand, within IAI47_04635 at 976.595 kb on + strand, within IAI47_04635 at 976.680 kb on + strand, within IAI47_04635 at 976.680 kb on + strand, within IAI47_04635 at 976.680 kb on + strand, within IAI47_04635 at 976.680 kb on + strand, within IAI47_04635 at 976.680 kb on + strand, within IAI47_04635 at 976.680 kb on + strand, within IAI47_04635 at 976.680 kb on + strand, within IAI47_04635 at 976.681 kb on - strand, within IAI47_04635 at 976.682 kb on + strand, within IAI47_04635 at 976.682 kb on + strand, within IAI47_04635 at 976.693 kb on + strand, within IAI47_04635 at 976.693 kb on + strand, within IAI47_04635 at 976.693 kb on + strand, within IAI47_04635 at 976.693 kb on + strand, within IAI47_04635 at 976.693 kb on + strand, within IAI47_04635 at 976.693 kb on + strand, within IAI47_04635 at 976.694 kb on - strand, within IAI47_04635 at 976.749 kb on + strand at 976.750 kb on - strand at 976.778 kb on + strand at 976.778 kb on + strand at 976.778 kb on + strand at 976.778 kb on + strand at 976.779 kb on - strand at 976.779 kb on - strand at 976.779 kb on - strand at 976.779 kb on - strand at 976.779 kb on - strand at 976.779 kb on - strand at 976.779 kb on - strand at 976.779 kb on - strand at 976.779 kb on - strand at 976.779 kb on - strand at 976.806 kb on - strand at 976.814 kb on + strand at 976.815 kb on - strand at 976.815 kb on - strand at 976.815 kb on - strand at 976.815 kb on - strand at 976.815 kb on - strand at 976.815 kb on - strand at 976.840 kb on + strand at 976.840 kb on + strand at 976.840 kb on + strand at 976.840 kb on + strand at 976.840 kb on + strand at 976.841 kb on - strand at 976.841 kb on - strand at 976.868 kb on + strand at 976.868 kb on + strand at 976.868 kb on + strand at 976.897 kb on + strand at 976.898 kb on - strand at 976.907 kb on - strand at 976.928 kb on + strand, within IAI47_04640 at 976.929 kb on - strand, within IAI47_04640 at 976.929 kb on - strand, within IAI47_04640 at 976.929 kb on - strand, within IAI47_04640 at 976.994 kb on + strand, within IAI47_04640 at 976.994 kb on + strand, within IAI47_04640 at 976.994 kb on + strand, within IAI47_04640 at 976.994 kb on + strand, within IAI47_04640 at 976.995 kb on - strand, within IAI47_04640 at 977.081 kb on + strand, within IAI47_04640 at 977.081 kb on + strand, within IAI47_04640 at 977.081 kb on + strand, within IAI47_04640 at 977.082 kb on - strand, within IAI47_04640 at 977.082 kb on - strand, within IAI47_04640 at 977.082 kb on - strand, within IAI47_04640 at 977.091 kb on - strand, within IAI47_04640 at 977.097 kb on + strand, within IAI47_04640 at 977.097 kb on + strand, within IAI47_04640 at 977.097 kb on + strand, within IAI47_04640 at 977.097 kb on + strand, within IAI47_04640 at 977.097 kb on + strand, within IAI47_04640 at 977.097 kb on + strand, within IAI47_04640 at 977.097 kb on + strand, within IAI47_04640 at 977.097 kb on + strand, within IAI47_04640 at 977.097 kb on + strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.098 kb on - strand, within IAI47_04640 at 977.129 kb on - strand, within IAI47_04640 at 977.143 kb on + strand, within IAI47_04640 at 977.143 kb on + strand, within IAI47_04640 at 977.143 kb on + strand, within IAI47_04640 at 977.143 kb on + strand, within IAI47_04640 at 977.143 kb on + strand, within IAI47_04640 at 977.144 kb on - strand, within IAI47_04640 at 977.184 kb on + strand, within IAI47_04640 at 977.184 kb on + strand, within IAI47_04640 at 977.184 kb on + strand, within IAI47_04640 at 977.185 kb on - strand, within IAI47_04640 at 977.185 kb on - strand, within IAI47_04640 at 977.185 kb on - strand, within IAI47_04640 at 977.185 kb on - strand, within IAI47_04640 at 977.185 kb on - strand, within IAI47_04640 at 977.209 kb on + strand, within IAI47_04640 at 977.210 kb on - strand, within IAI47_04640 at 977.240 kb on - strand, within IAI47_04640 at 977.256 kb on + strand, within IAI47_04640 at 977.256 kb on + strand, within IAI47_04640 at 977.256 kb on + strand, within IAI47_04640 at 977.256 kb on + strand, within IAI47_04640 at 977.257 kb on - strand, within IAI47_04640 at 977.257 kb on - strand, within IAI47_04640 at 977.257 kb on - strand, within IAI47_04640 at 977.257 kb on - strand, within IAI47_04640 at 977.321 kb on + strand, within IAI47_04640 at 977.322 kb on - strand, within IAI47_04640 at 977.341 kb on + strand, within IAI47_04640 at 977.353 kb on + strand, within IAI47_04640 at 977.354 kb on - strand, within IAI47_04640 at 977.373 kb on - strand, within IAI47_04640 at 977.419 kb on + strand, within IAI47_04640 at 977.420 kb on - strand, within IAI47_04640 at 977.420 kb on - strand, within IAI47_04640 at 977.486 kb on + strand, within IAI47_04640 at 977.501 kb on + strand, within IAI47_04640 at 977.513 kb on + strand, within IAI47_04640 at 977.702 kb on + strand, within IAI47_04640 at 977.702 kb on + strand, within IAI47_04640 at 977.702 kb on + strand, within IAI47_04640 at 977.702 kb on + strand, within IAI47_04640 at 977.702 kb on + strand, within IAI47_04640 at 977.703 kb on - strand, within IAI47_04640 at 977.703 kb on - strand, within IAI47_04640 at 977.703 kb on - strand, within IAI47_04640 at 977.703 kb on - strand, within IAI47_04640 at 977.731 kb on + strand, within IAI47_04640 at 977.732 kb on - strand, within IAI47_04640 at 977.737 kb on + strand, within IAI47_04640 at 977.738 kb on - strand, within IAI47_04640 at 977.738 kb on - strand, within IAI47_04640 at 977.779 kb on + strand at 977.780 kb on - strand at 977.780 kb on - strand at 977.780 kb on - strand at 977.913 kb on + strand at 977.913 kb on + strand at 977.960 kb on - strand at 977.960 kb on - strand at 977.960 kb on - strand at 977.970 kb on + strand at 977.970 kb on + strand at 977.971 kb on - strand at 977.971 kb on - strand at 978.043 kb on + strand, within IAI47_04645 at 978.043 kb on + strand, within IAI47_04645 at 978.150 kb on + strand, within IAI47_04645 at 978.150 kb on + strand, within IAI47_04645 at 978.151 kb on - strand, within IAI47_04645 at 978.151 kb on - strand, within IAI47_04645 at 978.151 kb on - strand, within IAI47_04645 at 978.151 kb on - strand, within IAI47_04645 at 978.158 kb on + strand, within IAI47_04645 at 978.158 kb on + strand, within IAI47_04645 at 978.198 kb on + strand, within IAI47_04645 at 978.198 kb on + strand, within IAI47_04645 at 978.229 kb on + strand, within IAI47_04645 at 978.229 kb on + strand, within IAI47_04645 at 978.230 kb on - strand, within IAI47_04645 at 978.230 kb on - strand, within IAI47_04645 at 978.230 kb on - strand, within IAI47_04645 at 978.230 kb on - strand, within IAI47_04645 at 978.270 kb on + strand, within IAI47_04645 at 978.270 kb on + strand, within IAI47_04645 at 978.270 kb on + strand, within IAI47_04645 at 978.270 kb on + strand, within IAI47_04645 at 978.271 kb on - strand, within IAI47_04645 at 978.271 kb on - strand, within IAI47_04645 at 978.314 kb on + strand, within IAI47_04645 at 978.426 kb on + strand, within IAI47_04645 at 978.426 kb on + strand, within IAI47_04645 at 978.426 kb on + strand, within IAI47_04645 at 978.539 kb on + strand, within IAI47_04645 at 978.540 kb on - strand, within IAI47_04645 at 978.630 kb on + strand, within IAI47_04645 at 978.630 kb on + strand, within IAI47_04645 at 978.630 kb on + strand, within IAI47_04645 at 978.630 kb on + strand, within IAI47_04645 at 978.631 kb on - strand, within IAI47_04645 at 978.726 kb on + strand, within IAI47_04645 at 978.726 kb on + strand, within IAI47_04645 at 978.726 kb on + strand, within IAI47_04645 at 978.726 kb on + strand, within IAI47_04645 at 978.726 kb on + strand, within IAI47_04645 at 978.726 kb on + strand, within IAI47_04645 at 978.726 kb on + strand, within IAI47_04645 at 978.726 kb on + strand, within IAI47_04645 at 978.727 kb on - strand, within IAI47_04645 at 978.727 kb on - strand, within IAI47_04645 at 978.727 kb on - strand, within IAI47_04645 at 978.727 kb on - strand, within IAI47_04645 at 978.727 kb on - strand, within IAI47_04645
Per-strain Table
Position Strand Gene LocusTag Fraction Survival; Treatment=UGA synthetic gw pH3.9 with 1 mM Glucose and 5 mM NaH2PO4; TreatmentDuration=24hrs; Outgrowth=LBkan50 remove 975,873 + IAI47_04630 0.72 -1.9 975,874 - IAI47_04630 0.72 -1.3 975,933 + IAI47_04630 0.79 +1.3 975,933 + IAI47_04630 0.79 +1.1 975,934 - IAI47_04630 0.79 +0.4 975,934 - IAI47_04630 0.79 -0.6 975,934 - IAI47_04630 0.79 +0.4 975,934 - IAI47_04630 0.79 -0.6 975,972 + IAI47_04630 0.83 -0.8 975,972 + IAI47_04630 0.83 -0.8 975,972 + IAI47_04630 0.83 -2.6 975,972 + IAI47_04630 0.83 +0.5 975,973 - IAI47_04630 0.83 -0.2 975,973 - IAI47_04630 0.83 -0.4 975,973 - IAI47_04630 0.83 +0.4 976,188 + +1.1 976,205 + IAI47_04635 0.12 +2.4 976,205 + IAI47_04635 0.12 +1.6 976,206 - IAI47_04635 0.13 +0.1 976,206 - IAI47_04635 0.13 -0.3 976,210 - IAI47_04635 0.13 -0.6 976,210 - IAI47_04635 0.13 +0.1 976,210 - IAI47_04635 0.13 +0.8 976,210 - IAI47_04635 0.13 -0.4 976,253 + IAI47_04635 0.19 +0.0 976,253 + IAI47_04635 0.19 -2.7 976,253 + IAI47_04635 0.19 +1.3 976,253 + IAI47_04635 0.19 +1.6 976,253 + IAI47_04635 0.19 -0.5 976,253 + IAI47_04635 0.19 +1.7 976,254 - IAI47_04635 0.19 +1.5 976,264 + IAI47_04635 0.21 +0.4 976,264 + IAI47_04635 0.21 +1.2 976,264 + IAI47_04635 0.21 +0.3 976,265 - IAI47_04635 0.21 -1.5 976,265 - IAI47_04635 0.21 +1.7 976,277 + IAI47_04635 0.23 +0.5 976,294 + IAI47_04635 0.25 +0.3 976,303 + IAI47_04635 0.27 -0.5 976,303 + IAI47_04635 0.27 +2.0 976,303 + IAI47_04635 0.27 -0.5 976,303 + IAI47_04635 0.27 -0.9 976,303 + IAI47_04635 0.27 +0.9 976,303 + IAI47_04635 0.27 +0.6 976,303 + IAI47_04635 0.27 +1.9 976,304 - IAI47_04635 0.27 +0.0 976,304 - IAI47_04635 0.27 -0.5 976,304 - IAI47_04635 0.27 +1.0 976,304 - IAI47_04635 0.27 +0.6 976,304 - IAI47_04635 0.27 -0.1 976,304 - IAI47_04635 0.27 -1.1 976,304 - IAI47_04635 0.27 +0.0 976,304 - IAI47_04635 0.27 -1.9 976,309 + IAI47_04635 0.27 -1.2 976,368 - IAI47_04635 0.36 -0.6 976,429 - IAI47_04635 0.45 +0.5 976,430 + IAI47_04635 0.45 +1.7 976,430 + IAI47_04635 0.45 +0.1 976,431 - IAI47_04635 0.45 -0.4 976,431 - IAI47_04635 0.45 +0.1 976,431 - IAI47_04635 0.45 -0.9 976,431 - IAI47_04635 0.45 -0.9 976,432 + IAI47_04635 0.45 +0.0 976,432 + IAI47_04635 0.45 +1.3 976,432 + IAI47_04635 0.45 +1.8 976,432 + IAI47_04635 0.45 -0.4 976,432 + IAI47_04635 0.45 -0.0 976,432 + IAI47_04635 0.45 +0.6 976,432 + IAI47_04635 0.45 +0.6 976,432 + IAI47_04635 0.45 -2.0 976,433 - IAI47_04635 0.45 -0.9 976,433 - IAI47_04635 0.45 -0.1 976,433 - IAI47_04635 0.45 +0.1 976,433 - IAI47_04635 0.45 -0.3 976,433 - IAI47_04635 0.45 +0.4 976,433 - IAI47_04635 0.45 +0.4 976,433 - IAI47_04635 0.45 +0.3 976,433 - IAI47_04635 0.45 -0.3 976,433 - IAI47_04635 0.45 +1.3 976,442 + IAI47_04635 0.47 -0.1 976,442 + IAI47_04635 0.47 -0.7 976,442 + IAI47_04635 0.47 -0.8 976,442 + IAI47_04635 0.47 -0.3 976,442 + IAI47_04635 0.47 +0.4 976,442 + IAI47_04635 0.47 -0.2 976,442 + IAI47_04635 0.47 +0.5 976,442 + IAI47_04635 0.47 -0.6 976,442 + IAI47_04635 0.47 +0.6 976,442 + IAI47_04635 0.47 -0.4 976,442 + IAI47_04635 0.47 -1.3 976,443 - IAI47_04635 0.47 +0.6 976,443 - IAI47_04635 0.47 -0.5 976,443 - IAI47_04635 0.47 +1.0 976,443 - IAI47_04635 0.47 -0.6 976,444 + IAI47_04635 0.47 -0.2 976,444 + IAI47_04635 0.47 -0.1 976,444 + IAI47_04635 0.47 +0.5 976,444 + IAI47_04635 0.47 -0.3 976,444 + IAI47_04635 0.47 +0.7 976,444 + IAI47_04635 0.47 -0.3 976,444 + IAI47_04635 0.47 +2.4 976,445 - IAI47_04635 0.47 -1.0 976,445 - 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Or see this region's nucleotide sequence