Pairwise Alignments

Query, 4679 a.a., peptide synthase from Pseudomonas simiae WCS417

Subject, 4317 a.a., non-ribosomal peptide synthase (subunit of ferribactin synthase) from Pseudomonas putida KT2440

 Score = 1933 bits (5007), Expect = 0.0
 Identities = 1179/2615 (45%), Positives = 1550/2615 (59%), Gaps = 116/2615 (4%)

Query: 45   QVEDRQA--LSYAQQRMWFLWQLDPASGAYNLPGAVRLKGALSLDALEQAFASLVARHET 102
            +V+ RQA  LSY+QQRMWFLWQ++P S AYN+ G  RL+G L +DA E+A  +L+ RHET
Sbjct: 1712 RVDRRQAVPLSYSQQRMWFLWQMEPDSPAYNVGGMARLRGTLHVDAFERALQALIVRHET 1771

Query: 103  LRTVFQRQADERLLQVAADTSLTVEQLDLSHLALAEREEAVNAAATRQSLLPFDLENGPL 162
            LRT F          VA D+ L ++  D S L+    ++ +   A  Q+  PFDLE GPL
Sbjct: 1772 LRTTFPSIDGVPYQCVAEDSGLHLDWQDFSALSDDACQQRLQQLADDQAHQPFDLERGPL 1831

Query: 163  LRVQLLKLDAQEHVLLLTLHHIVSDGWSMNVLIDEFIRCYDAHERSQAPQLPALPIQYSD 222
            LR  L+K + +EH  +LTLHHIV++GW+M++   E    Y+A    +   L  LP+QY D
Sbjct: 1832 LRACLVKANEREHFFVLTLHHIVTEGWAMDIFARELGELYEAFVDDRESPLAPLPVQYLD 1891

Query: 223  YALWQRRWLEAGEQARQLEYWQARLGDEHPVLELPTDRPRPAMPSYQGTRHDFAINPQLA 282
            Y++WQR+WLE+GE ARQL YW+ RLGDEHPVL LP DRPRPA+ S++G  + F ++P L 
Sbjct: 1892 YSVWQRQWLESGEGARQLAYWKHRLGDEHPVLALPADRPRPAVQSHRGELYRFELDPALV 1951

Query: 283  AQLRTCAQKHNVTLFMLLLGAFNVLLHRYTGQGDIRVGVPIANRNRTEVEGLIGFFVNTQ 342
            A++     +  +TLFM +      LLHRY+GQ D+R+G P+ANR R E EGLIG F+NTQ
Sbjct: 1952 ARVHAFNSQRGLTLFMTMTATLAALLHRYSGQRDLRIGAPVANRIRPESEGLIGAFLNTQ 2011

Query: 343  VLRTELTGQTLVTELLQSIKEHALGAQAHQELPFERLVEALKVERSLSHTPLFQVMYNHQ 402
            VLR EL GQ   + LL+ +++ A+  Q+HQ+LPF++LVEAL+  RS ++ PLFQVM N Q
Sbjct: 2012 VLRCELDGQMTASALLEQVRQAAIEGQSHQDLPFDQLVEALQPPRSSAYNPLFQVMCNVQ 2071

Query: 403  PVVADIANVSTASGLELALVEWQGRTTQFDLTLDTYEKSGTLHAALTYANDLFDALTIER 462
                  +   T +G+++  +      T+FDL L+  +  G L   LTY+ DLFDA  I R
Sbjct: 2072 RWAFQQSR--TLAGMQVDYLVNDASATKFDLYLEVTDLDGRLGCCLTYSRDLFDAPRIAR 2129

Query: 463  MAGHWIRLLQAMVADGTQRIGELPMLAADEQQLLVHAWNQTAEAYPTERGIHHLVEDQVQ 522
            MA HW +LL  ++ +   R+ ELPML++ EQQ+L     Q    +   + +H L   Q  
Sbjct: 2130 MAEHWQQLLVGLLDNPQHRLCELPMLSSAEQQVLTGQL-QGEHDFDLGQTLHGLFAAQAA 2188

Query: 523  ATPDAPALVFGTTTLTYAQLDARANQLAHALREKGVAPDVLVGICIERSIEMVVGLLAIL 582
             TP A AL F    LTYA+LD +AN+LA ALRE+GV P V VG+ +ERS+EMVVGLLAIL
Sbjct: 2189 RTPQAGALTFAGQHLTYAELDQQANRLARALRERGVGPQVRVGLALERSLEMVVGLLAIL 2248

Query: 583  KAGGAYVPLDPEYPQERLAYMIEDSGIQLLLSQHSLLPVLPV--EGIQVIALDQPASWLD 640
            KA GAYVPLDPEYP +RL YMIEDS I LLLSQ +LL  L    EG+   +L+  A+ L 
Sbjct: 2249 KADGAYVPLDPEYPLDRLRYMIEDSRIGLLLSQRALLHTLGELPEGVASWSLEDDAASLA 2308

Query: 641  GYS-AESANVSLHALNLAYVIYTSGSTGKPKGAGNSHRALVNRLCWMQQAYGLDARDAVL 699
             YS A   N+SL   + AY+IYTSGSTGKPKG   SH         +  A+G+ + D  L
Sbjct: 2309 AYSDAPLDNLSLPQ-HQAYLIYTSGSTGKPKGVVVSHGEFAMHCQAVIAAFGMRSDDCEL 2367

Query: 700  QKTPFSFDVSVWEFFWPLMTGARLVVAAPGEHREPARLIETIGRHGITTLHFVPS---ML 756
                 +FD +    + PL+ GAR+V+ A G+      + + +    ++ L F PS    L
Sbjct: 2368 HFYSINFDAASERLWVPLLCGARVVLRAQGQWGAED-ICQLVREQQVSILGFTPSYGSQL 2426

Query: 757  QAFIHEPGVQACTSLQRIVCSGEALPLDAQLQVFAKLPQAGLFNLYGPTEAAIDVTHWTC 816
              ++   G Q    L  ++  GEAL  +   ++         FN YGPTE    V    C
Sbjct: 2427 AQYLGGRGEQLPVRL--VITGGEALTGEHLQRIRQAFAPQQCFNAYGPTETV--VMPLAC 2482

Query: 817  VDEGA-----DSVPIGRPIANLGTYVLDAQLNPVPAGVSGELYLGGIGLARSYHRRPALT 871
            +   A      SVPIGR I +   Y+LD  L  +P G  GELY+GG GLA+ YH RP LT
Sbjct: 2483 LAPEALPADLGSVPIGRVIGSRTAYILDEDLALLPQGGIGELYVGGAGLAQGYHDRPGLT 2542

Query: 872  AERFVPSPFG-DGARLYRTGDRVRQRADGVIEYLGRLDHQVKLRGLRIELGEIETRLMQH 930
            AERFV  PF  +G RLYRTGD VR   DG++EY+GR D QVK+RG RIELGEIE+RL+ H
Sbjct: 2543 AERFVADPFSAEGGRLYRTGDLVRLGTDGLVEYVGRADQQVKIRGFRIELGEIESRLLAH 2602

Query: 931  PSVREAVVL---VQGGKQLVAYLVLED-TAPVDLKAWLLSS--------LPEYMVPSHMV 978
              V EAVVL   + GGKQLV YLV    TA  D++  L  +        LP+YMVP+H+V
Sbjct: 2603 EDVDEAVVLALDLPGGKQLVGYLVCRQATAGNDVQQLLREAVKADARQHLPDYMVPAHLV 2662

Query: 979  PLAKLPVTANGKLDRKALPLPDVAP-QQAFVAPENDLQKAVAAIWSDVLGVEQVGLDDNF 1037
             L  LP+  NGKLDR ALPLPD+   +Q + AP N+++  +A IW DVL V +VG+ DNF
Sbjct: 2663 LLDSLPLMGNGKLDRHALPLPDLEQARQQYQAPGNEVEAQLAQIWRDVLNVARVGVQDNF 2722

Query: 1038 FELGGDSIISIQVVSRARQAGIRLSPRDLFQYQSVRSLALVATF-EQATAIDQGPVTGEV 1096
            FELGGDSI+SIQVVSRARQAG++ +PRDLFQ+Q++++LA V    E A+ ++QG   G+ 
Sbjct: 2723 FELGGDSILSIQVVSRARQAGLQFTPRDLFQHQTIQTLATVVQLSEAASTVEQGLRQGQA 2782

Query: 1097 ILTPVQHSFFEQAIPARQHWNQSLLLTPREALEPARLEAALTRLINHHDALRLRFLQQAD 1156
             LTP+QH FF+  +P  QHWNQ++LL  R+ L  A LE AL  L+ HHD+LRLRF     
Sbjct: 2783 GLTPIQHWFFDSDVPQPQHWNQTVLLEVRQPLVEATLEQALAALVQHHDSLRLRFSDAQG 2842

Query: 1157 GWRQAHAEPVATASLWQSQVADEAELAALCDEAQRSLDLEQGPLLRAVLAVMADGTQRLL 1216
             W+  + +P A   LW + VAD  +  AL  + QRSLDL+QGPLLRA+L     GTQRLL
Sbjct: 2843 RWQAQYVQPGAEQLLWSATVADFNDCQALYTDVQRSLDLQQGPLLRALLVRDGQGTQRLL 2902

Query: 1217 LVVHHLAVDGVSWRILLEDLQQAYRNAALPAKTSAYQLWAQQLQRHA--QTLDEQLPYWQ 1274
            L +HHL VDGVSWR+LLEDLQ  YR   LPAKT A   WA +L  +A   +L ++L +WQ
Sbjct: 2903 LAIHHLVVDGVSWRVLLEDLQALYRGQPLPAKTHAVGDWAARLASYAGSDSLRDELAWWQ 2962

Query: 1275 AQTASA--DLPCDNPQGGLQNRLGSLLEIRLSAEHTRQLLQDAPAAYRTQVNDLLLTALA 1332
             Q      +LPCD+PQG   +R    L I L  E TRQLLQ APAAY TQVNDLLLTALA
Sbjct: 2963 GQLGGVRRELPCDHPQGSNLHRHAQTLAIGLDVEQTRQLLQLAPAAYHTQVNDLLLTALA 3022

Query: 1333 RVISRWSEQPAALIQLEGHGREDLFDSVDLSRTVGWFTSLFPVRLHA-----DGELSAAI 1387
            R + RWS     L+QLEGHGR+ LF+ +DL+R+VGWFT+ +P+ L       D   + +I
Sbjct: 3023 RALCRWSGDEEVLVQLEGHGRDGLFEDMDLTRSVGWFTNAYPLSLRPLPGEDDAARAGSI 3082

Query: 1388 KSVKEQLRAVPHKGIGYGLLRYLGTPDAREALSALAAPRITFNYLGQFDRQFNESALFVP 1447
            K +KEQLR VPHKG+GYG+LRYL     RE ++AL   RITFNYLGQFD+QF+ +ALF P
Sbjct: 3083 KRIKEQLRHVPHKGLGYGVLRYLADAAGREQMAALPQARITFNYLGQFDQQFDSAALFQP 3142

Query: 1448 ATQGSGQAQDPQAPLANWLTVEGQVYGGELALQWGFSREMFDVATVQRLADDYRAELQAL 1507
                +G A D  APL NWL+V+GQVYGG L L+W FS E +D  T+ RLA+ YR EL AL
Sbjct: 3143 LEASAGLAHDLDAPLPNWLSVDGQVYGGALQLRWTFSAERYDQQTIARLAEGYRQELLAL 3202

Query: 1508 IEHCCATPAGQVTPSDFPLARLTQQQLDGLPIAGPAIADLYPLSPMQQGMLFHTLYEPEA 1567
            + HC A   G  TPSDFPLA LTQ+Q+D LP+    I D+YPL+PMQ+GML HTL EP  
Sbjct: 3203 VAHCLADGNGSFTPSDFPLAHLTQEQIDALPVPAAQIEDVYPLTPMQEGMLLHTLLEPGT 3262

Query: 1568 EAYINQLRLDI-EGLDLLAFGRAWQAALDRHDILRSSFHWLGLDSAHQVIQR----QVDV 1622
              Y  Q R  I   LD   F +AWQA + RH+ LR+SF W   ++  Q+I +     VD 
Sbjct: 3263 GLYYMQDRYRINSALDPERFAQAWQAVVARHEALRASFSWNSGEAMLQIIHKPGNLAVDY 3322

Query: 1623 Q--LHVIEDI-DADCDALAVAEREKGFTLNAAPLFRLMLVRGAGTAWHLIFTSHHILMDG 1679
            Q    + +D  +    AL   ERE GF L +   F L LVR A   +  + ++HHIL+D 
Sbjct: 3323 QDWRGLADDAQEQRLQALHKQEREAGFALLSEAPFHLRLVRVAEERYWFMMSNHHILIDA 3382

Query: 1680 WSNAQLLGEVIAHY----AGHAVPAPL-GQFRDYLGWLHQQA--SGEAFWKATLAPLQTP 1732
            W  + L+ +    Y     G     P+  ++RDY+GWL +Q      A+W+A LA  +  
Sbjct: 3383 WCRSLLMNDFFEVYQALGEGRQAQLPVPPRYRDYIGWLQRQGLDDARAWWQANLAGFERA 3442

Query: 1733 TLLA--QALRVPTEGKGMAEHRVLLGDDFTR-------ALGEFARRHKVTLNTVLQGAWS 1783
            T +   + LR    G GM     ++GD +TR        L E A+ H++T+NT  Q AW+
Sbjct: 3443 TAIPSDRPLRHDHAGSGM-----VVGDCYTRLEMNDGVRLRELAQAHQLTVNTFAQAAWA 3497

Query: 1784 LLLQRYTGQDCVAFGATVAGRSAPLPGIEQQLGLFINTLPI---VSAASPALSVAGWLSE 1840
            L+L RY+G   VAFG TVAGR   +P +++ +GLFIN++ +   + AAS   SV  WL  
Sbjct: 3498 LVLSRYSGDRDVAFGVTVAGRPVSMPQMQRTVGLFINSVALRVQLPAASERCSVRQWLQG 3557

Query: 1841 LQALNLSLRDHEHVPLYDIQGWAG-QQGAALFDTLLVFENFPVAEAL---KQGAPAGLTF 1896
            L   N+ LR++E++PL  IQ  +   +G  LFD+L VFEN PV  A+    Q   A    
Sbjct: 3558 LLQSNMELREYEYLPLVAIQACSELPKGQPLFDSLFVFENAPVETAVLNHAQHLNASSDS 3617

Query: 1897 GRLHNHEQTHYPLTLGIELGASLRLEFGYDQARFSAQQVARLSGNLHHLLVQMLADAQAP 1956
            GR H    T++PLT     G  L L   +DQ  F    V RL      LL+ ++   +  
Sbjct: 3618 GRTH----TNFPLTAVCYPGDDLGLHLSFDQRYFDYPTVERLLAEFKRLLLALVQGFEGE 3673

Query: 1957 LGNLRLLDAATHRDVLAH-SRATGAAPRQLRVHERIAAQAAATPDALAVQAGDARLSYAQ 2015
            +  L LL A     +L   +R     P +     +  AQ  A P     +  +A   YA 
Sbjct: 3674 VSELPLLSAQEQGFLLEDCNRTERVYPLEQSYIAQFEAQVVAHPQRTVARCLEASYDYAG 3733

Query: 2016 LNDHANRLAHRLLALGVGPGQRVGLAARRGPQLIVSLLAVLKSGAAYVPLDPNYPTERLA 2075
            LN  ANRL H L+A GV   Q V L A RG  L+  ++   K+GA Y+PLDP  P  RL 
Sbjct: 3734 LNLAANRLGHALVAAGVCVDQPVALLAERGLPLLGMIVGSFKAGAGYLPLDPGLPAARLQ 3793

Query: 2076 YMLADSRLDLLL-----SETG--LLADLPL---PQGLARLDFTASGAELADYPSTPPANH 2125
             ++  SR  +L+     +E G  LL +L     PQ L   D  A+        S  P  H
Sbjct: 3794 SIIQLSRTPVLVCSAACAEQGRQLLGELEASLRPQLLVWEDVQAN-----PIASHNPGIH 3848

Query: 2126 AAAADLAYVIYTSGSTGLPKGVAIDHAALSQFCDSAEAYSRLSASDRVLQFATFSFDGFV 2185
            +   +LAYVIYTSGSTGLPKGV ++   +     S   Y  LS  D + Q A+ SFD  V
Sbjct: 3849 SGPDNLAYVIYTSGSTGLPKGVMVEQRGMLNNQLSKVPYLALSEQDVIAQTASQSFDISV 3908

Query: 2186 EQCYPPLCVGAAL-IMRGDELWDAGQLAREIVEQGVT-LADLPAAYWYLLAKECAVDRRT 2243
             Q       GA + I+      D   L   +   G+T L  +P+    +LA     D + 
Sbjct: 3909 WQFLAAPLFGAKVEIVPNVIAHDPQGLLAHVQATGITVLESVPSLIQGMLAS----DHQA 3964

Query: 2244 LGNLRQVHVGGEAMSVEGLRAWHTAGLGGVRLVNTYGPTEATVVSSVHDCQLADASDAFG 2303
            L  LR +   GE M  E    W       + LVN YGP E +   +      A    ++ 
Sbjct: 3965 LDGLRWMLPTGEVMPPELAAQW-LQRYPQIGLVNAYGPAECSDDVAFFRVDAASTQGSY- 4022

Query: 2304 VPIGRAIEGRALYVLDSGFELLASDGVGELCIGAEHGLAQCYFDRPALTAERFLPDPFSA 2363
            +PIG   +   LY+      L+    VGELC+ A  G+ + Y   P  TA  F+P P+ A
Sbjct: 4023 LPIGTPTDNNRLYLFGEDQALVPLGAVGELCV-AGAGVGRGYVGDPVRTALAFIPHPYGA 4081

Query: 2364 VPGARLYRSGDLARYNEAGALEYVGRIDHQVKIRGFRVEMGEIEASLQALPSLREAAVIA 2423
             PG RLYR+GDLAR    G LEYVGRIDHQVKIRG+R+E+GEIEA L     +R+AAV  
Sbjct: 4082 -PGERLYRTGDLARQRLDGVLEYVGRIDHQVKIRGYRIELGEIEARLHEQAEIRDAAVAV 4140

Query: 2424 QPSATGTQLVAYVVP---VTAQTLDPQALAATLRQSLPDYMVPGQWVILEALPLNNNGKL 2480
            Q    G  LV Y+V    V+A     + +   LR  LP+YMVP  W   ++LP N NGKL
Sbjct: 4141 QEGVNGKHLVGYLVAHQGVSADAALLEQVKQRLRAELPEYMVPLHWGWFDSLPHNANGKL 4200

Query: 2481 DRRALPAPDL-NQARQAYLAPQNPLQSRLAAIWQAVLQVEQVGLNDHFFERGGHSLLATQ 2539
            DR+ALPA D+  Q  QAYLAP+N L+  LA IW  VL+ E+VG++D+FFE GGHSLLATQ
Sbjct: 4201 DRKALPAIDIGGQYSQAYLAPRNELEDVLAGIWADVLKAERVGVHDNFFELGGHSLLATQ 4260

Query: 2540 VVSRVRHDLKLEVPLRALFEQPTLEAFAAACAGLQ 2574
            + SRV+  L+L VPLRA+FE  T+EA A    GLQ
Sbjct: 4261 IASRVQKQLQLNVPLRAMFECSTVEALAEYVQGLQ 4295



 Score = 1789 bits (4633), Expect = 0.0
 Identities = 1154/2804 (41%), Positives = 1560/2804 (55%), Gaps = 149/2804 (5%)

Query: 1978 TGAAPRQLRVHERIAAQAAATPDALAVQ------AGDARLSYAQLNDHANRLAHRLLALG 2031
            T A    L + + +A +AA TP+ +A++         A LSY  L+  A  +A  L    
Sbjct: 2    TDAFELPLSLVQALAQRAAQTPERIALRFLADTPGEQAVLSYRDLDQRARTIAAALQGRA 61

Query: 2032 VGPGQRVGLAARRGPQLIVSLLAVLKSGAAYVPLDP-----NYPTERLAYMLADSRLDLL 2086
               G R  L    GP  + +    L +G   VP  P         ERL  ++ D+   LL
Sbjct: 62   -SFGDRAVLLFPSGPDYVAAFFGCLYAGVIAVPAYPPESARQQHQERLLSIIDDAEPRLL 120

Query: 2087 LSETGLLADLPLPQGLARLDFTASGAELADYPSTPPANHA-------AAADLAYVIYTSG 2139
            L+   L   L   QGL  L    +  +L       PA  A          D+A++ YTSG
Sbjct: 121  LTVAALRDSL---QGLEALA-ADNAPQLLAVDGLDPALAANWREPVLKGDDIAFLQYTSG 176

Query: 2140 STGLPKGVAIDHAAL---SQFCDSAEAYSRLSASDRVLQFATFSFD-GFVEQCYPPLCVG 2195
            ST LPKGV + H  L    Q          L+  D ++ +     D G +     P+  G
Sbjct: 177  STALPKGVQVSHGNLVANEQLIRQGFGID-LNPDDVIVSWLPLYHDMGLIGGLLQPIFSG 235

Query: 2196 AALIMRGDELWDAGQLA--REIVEQGVTLADLPAAYWYLLAK---ECAVDRRTLGNLRQV 2250
               ++     + A  L   + I E G T++  P   + L ++   E ++    L   R  
Sbjct: 236  VPCVLMSPRYFLARPLRWLQAISEYGGTISGGPDFAYRLCSERVSEASLAGLDLSRWRVA 295

Query: 2251 HVGGEAMSVEGLRAW----HTAGLGGVRLVNTYGPTEATV-VSSVHDCQ--LADASDAFG 2303
            + G E +  + L  +       G        +YG  EAT+ VS     Q   A   DA  
Sbjct: 296  YSGSEPIRQDSLATFADKFQACGFDPQSFFASYGLAEATLFVSGSRRGQGIAALELDAKA 355

Query: 2304 VPIGRAIEGRALYVLDSGF---------------ELLASDGVGELCIGAEHGLAQCYFDR 2348
                RA  G    ++  GF               ++L  + VGE+  G    +A+ Y+  
Sbjct: 356  FAANRAEPGTGSVLMSCGFPQPGHAVRIVEPQQLQVLGDNQVGEIWAGGP-SIARGYWRN 414

Query: 2349 PALTAERFLPDPFSAVPGARLYRSGDLARYNEAGALEYVGRIDHQVKIRGFRVEMGEIEA 2408
            P  +A  F+      + G    R+GDL    E G +   GR+   + +RG  +   ++E 
Sbjct: 415  PEASARTFVE-----MDGQTWLRTGDLGFIRE-GEVFVTGRLKDMLIVRGQNLYPQDLEK 468

Query: 2409 SLQA-LPSLREAAV----IAQPSATGTQLVAYVVPVTAQTLDPQALAATLRQSLPDYMVP 2463
            +L+  +  LR+  V    +      G  +   +     + L PQ L  TLRQ + D    
Sbjct: 469  ALEREVQVLRKGRVAVFAVEHQGEEGIGVAVEISRNVQKALKPQDLIKTLRQVVADACRQ 528

Query: 2464 GQWVIL----EALPLNNNGKLDRRALPA-------------PDLNQARQAYLAPQNPLQS 2506
               V+L     ALP  ++GKL R A                P+++ A     A    LQ+
Sbjct: 529  APAVVLLLNPGALPKTSSGKLQRSACRLRMDDGSLDCYARFPEVSAAGIRATAGDE-LQA 587

Query: 2507 RLAAIWQAVLQVEQVGLNDHFFERGGHSLLATQVVSRVRHDLKLEVPLRALFEQPTLEAF 2566
            R+AA+W+ +L+VE V  +DHF   GG+S+ ATQ  +R+  +L +++ LR LFE P L  +
Sbjct: 588  RIAAVWRDILKVEAVAADDHFLLLGGNSIAATQATARLADELGIDLSLRTLFEAPVLADY 647

Query: 2567 AAACAGLQTAPALPLVAVT---RDQPLALSFAQERQWFLWQLDPTSAAYHVPSALHLRGR 2623
            + A A +    A    A+T   R Q L  S AQ R W LWQL+P SAAY++P+ LHLRG 
Sbjct: 648  SKAVAQIIADGAAATTAITTLERAQALPQSLAQNRLWLLWQLEPQSAAYNIPAGLHLRGE 707

Query: 2624 LDVAALEQAFQVLVQRHEPLRTTFVEEGEHTWQVIHPRLAMPIERQRVVASAIEHAVAQ- 2682
            LDV AL  AFQ LV RHE LRT F EE     Q I P+  + + R  +   + E   AQ 
Sbjct: 708  LDVNALHVAFQALVARHESLRTVFSEENGQALQRILPQQPLSLHRLDLEGHSPEQVAAQR 767

Query: 2683 --EIQRPFDLVNGPLMRVNLLEVEAEHHVLVITQHHIISDGWSMQVMVDELVALY----Q 2736
              E ++PFDL  GPL+RV L+ +  E H L +T HHI++DGWS+ ++++E   LY    Q
Sbjct: 768  ETEARQPFDLTQGPLLRVTLVRLGDEEHQLWVTLHHIVADGWSLNILLEEFAKLYAARCQ 827

Query: 2737 G-QAELPALPIQYVDYAQWQRDWMAAGEKQRQLDYWRTRLGTEHPVLELPLDHPRPALQS 2795
            G +A L  LP+ Y DY  WQR W+A GE +RQL YW+  LG E P L+L  DHPR + + 
Sbjct: 828  GLEANLAPLPLGYADYGSWQRQWLADGEAERQLQYWKAHLGGELPALDLCTDHPRASQRE 887

Query: 2796 HRGARRQVQLDNALVAELKALAQRQEVTLFVLLLASFQTLLHRYSGQSRIRVGVPIANRN 2855
            H  AR  +++   L   LK LA +Q+ +LF++LLA +Q LL RYSGQ+ IRVGVP ANR 
Sbjct: 888  HSAARFSLKVPVRLGEALKGLAHQQQASLFMVLLAGWQALLQRYSGQADIRVGVPNANRP 947

Query: 2856 RLETERLLGFFVNTQVLQADVHGQMAFDQLLAQVKQQALDAQAHQDLPFEQLVEALQPER 2915
            RLET+ ++GFF+NTQVL+A   G+ +F QLL QV+Q  L AQA+QDLPFEQLVEAL   R
Sbjct: 948  RLETQGMVGFFINTQVLRARFDGRHSFTQLLTQVRQATLQAQANQDLPFEQLVEALPDAR 1007

Query: 2916 SLSHNPLFQVMFNHQDNLRAAPLQLPGLDLQAVDWAGHSTQFDLNLETEESAEG-LWASL 2974
                  LFQVMFNHQ    +A  +LPGL  + + W     +FDL L +EE   G L  + 
Sbjct: 1008 EQG---LFQVMFNHQQRDLSALRRLPGLLAEELAWHSREAKFDLQLHSEEDHLGRLSLAF 1064

Query: 2975 TYATDLFDATTVERLAEHWQNLLRAVVNDASQAVDELAMLSAPQWQQMVEAWNDTSVDYP 3034
             YA +LF+A+TVERLA H   LL  V     QA+ ++ +L  P   Q++  W        
Sbjct: 1065 DYAAELFEASTVERLAHHLLALLEQVCAAPQQALGDVQLLDEPGRAQLL-GWGQAPAAAA 1123

Query: 3035 RERCVHTLFEARALAQPDALALQFKGQALSYGELNRRANRLAHRLIAAGVGPDVLVAVHV 3094
            +   V  L E   L  P   AL + G +L Y  L+++ANRLAH L   GVGPD  VA+ +
Sbjct: 1124 QHLLVEQLNEQVRLT-PQRTALVWDGGSLDYAGLHQQANRLAHYLRDKGVGPDTCVAIAI 1182

Query: 3095 ERSLDMVIGLLATLKAGGAYVPLDPQFPADRLAFMLEDSRARVLLTQSHLDGSLAHPQGV 3154
            ERS  +++GLLA LKAGGAYVPLD  +PA+RLA+ML D  AR+LL+ S L G L    GV
Sbjct: 1183 ERSPQLLVGLLAILKAGGAYVPLDVDYPAERLAYMLADCDARLLLSHSSLLGKLPQVDGV 1242

Query: 3155 QVLMVEAD--DNAVEHDPQVNVTPEHLAYVIYTSGSTGTPKGVMVRHEALCSFTCGMAGT 3212
              + ++    D+   H P +++  ++LAYVIYTSGSTG PKGV   H AL      M  T
Sbjct: 1243 SAIALDQLHLDSWPSHAPGLHLHGDNLAYVIYTSGSTGQPKGVGNTHAALAERLQWMQTT 1302

Query: 3213 LDIGADARVLSLTTFSFDIFALELYVPLTVGATVLLSGQELALDPEAIIDLVHSQCANVL 3272
              +     ++     SFD+   E + PL  G  ++L+G     DP+ I  LV +     L
Sbjct: 1303 YALNDSDVLMQKAPISFDVSVWECFWPLVTGCKLVLAGPGEHRDPQRIAALVQAYGVTTL 1362

Query: 3273 QATPSTWRMLLESPRAQLLRGIKCL-CGGEALPSDLAQRLLDL--QGPLWNLYGPTETTI 3329
               P   ++ ++ P+A     ++ +  GGEAL + L  R+L +  Q  L N YGPTET I
Sbjct: 1363 HFVPPLLQVFVQEPQASACSSLRWVFSGGEALSATLRDRVLQVLPQVQLHNRYGPTETAI 1422

Query: 3330 ----WSAAHRLHEAQPLVGRPIANTALFILNAGLTPSPVGAAGELLIGGVGLARGYHERP 3385
                W       E  P +GRP+ N    +L+  L  S  G  GEL +GG GLARGY  RP
Sbjct: 1423 NVTHWHCQVVDGERSP-IGRPLGNVLCRVLDDELELSAPGVPGELYLGGAGLARGYLGRP 1481

Query: 3386 ALTAERFVPNPYGAPGERLYRTGDLARYRAD-GVVEYIGRVDHQVKVRGFRIELGEVEAC 3444
             LTAERFVP   G  G+RLYR+GD AR++     +EY+GR+D QVKVRGFR+E  EV+AC
Sbjct: 1482 GLTAERFVPQADGN-GQRLYRSGDRARWQVQLEALEYLGRLDQQVKVRGFRVEPEEVQAC 1540

Query: 3445 LREQAGIREAVVLADNDRLIAYLVS--SAPDAPEVYKAALRERLPDYMVPAHLMFLDSLP 3502
            L  QAG+ +A+VL   D + A LV   S    P    A L E+LP YMVPA L+ L  +P
Sbjct: 1541 LLAQAGVEQALVLIHKDAIGAQLVGYYSGSAQPAEVLAVLAEQLPAYMVPAQLIPLAQMP 1600

Query: 3503 LTPNGKLDRKALPAVDAALLRTGHVAPVTPREQQVAAIWADVLDLPQVGLDDHFFELGGH 3562
            L P+GK+DRKALPA      +  HV P T  +QQVAAIW DVL+LP++GL D FF LGGH
Sbjct: 1601 LGPSGKVDRKALPA--PVWQQREHVEPRTELQQQVAAIWRDVLNLPRIGLQDDFFALGGH 1658

Query: 3563 SLLATRVVSRVRQALALEVALKTLFEHPLLGDF------VQALGEEGVVAPALLKADRRQ 3616
            SLLAT++VSR RQA  +E+ LK LFE   LG F      ++A GE  +   A+ + DRRQ
Sbjct: 1659 SLLATQIVSRTRQACDVELPLKALFEASELGAFCAEIARLRAAGERNLQG-AITRVDRRQ 1717

Query: 3617 PLPLSYAQERQWFLWQLDPHSTAYHIPSALRLHGQLDLGALQRSFDTLLARHESLRTHVR 3676
             +PLSY+Q+R WFLWQ++P S AY++    RL G L + A +R+   L+ RHE+LRT   
Sbjct: 1718 AVPLSYSQQRMWFLWQMEPDSPAYNVGGMARLRGTLHVDAFERALQALIVRHETLRTTFP 1777

Query: 3677 QDAAATVQ-VIEDSGLIEIDLAD----TDEAS---LKARVAEVVAQPFELLRGPLLRVKL 3728
                   Q V EDSGL  +D  D    +D+A    L+    +   QPF+L RGPLLR  L
Sbjct: 1778 SIDGVPYQCVAEDSGL-HLDWQDFSALSDDACQQRLQQLADDQAHQPFDLERGPLLRACL 1836

Query: 3729 LRLAADDHVLVLVQHHIVSDGWSMQLMVEELVQLYAAFSQGHMPDLPALPIQYADYALWQ 3788
            ++    +H  VL  HHIV++GW+M +   EL +LY AF       L  LP+QY DY++WQ
Sbjct: 1837 VKANEREHFFVLTLHHIVTEGWAMDIFARELGELYEAFVDDRESPLAPLPVQYLDYSVWQ 1896

Query: 3789 RNWMEAGEKARQLAYWQAQLGGVQPVLELPFDYLRPAVQSHRGARLGIELQPQLLAGLRR 3848
            R W+E+GE ARQLAYW+ +LG   PVL LP D  RPAVQSHRG     EL P L+A +  
Sbjct: 1897 RQWLESGEGARQLAYWKHRLGDEHPVLALPADRPRPAVQSHRGELYRFELDPALVARVHA 1956

Query: 3849 LAQGAGVTLPMVLLASYQALLHRYSGQEDVRVGVPIANRNRLETEGLIGFFVNTQVLKAD 3908
                 G+TL M + A+  ALLHRYSGQ D+R+G P+ANR R E+EGLIG F+NTQVL+ +
Sbjct: 1957 FNSQRGLTLFMTMTATLAALLHRYSGQRDLRIGAPVANRIRPESEGLIGAFLNTQVLRCE 2016

Query: 3909 IHGQMSVVQLLQQVRQRSLEAQAHQDLPFEQLVEALQPERSMSLSPLFQVLFNHRVTSAA 3968
            + GQM+   LL+QVRQ ++E Q+HQDLPF+QLVEALQP RS + +PLFQV+ N +   A 
Sbjct: 2017 LDGQMTASALLEQVRQAAIEGQSHQDLPFDQLVEALQPPRSSAYNPLFQVMCNVQ-RWAF 2075

Query: 3969 HHLQRLTELDVEVLSWDEGVAQFDLALDVEESQATLRASLNYATDLFAPATIERMAGHWQ 4028
               + L  + V+ L  D    +FDL L+V +    L   L Y+ DLF    I RMA HWQ
Sbjct: 2076 QQSRTLAGMQVDYLVNDASATKFDLYLEVTDLDGRLGCCLTYSRDLFDAPRIARMAEHWQ 2135

Query: 4029 NLLQAMVADQQQPISQLNLLGQDEQHHILQLWDQTDAGFSAKRLVHELVADRARENPDAV 4088
             LL  ++ + Q  + +L +L   EQ  +L    Q +  F   + +H L A +A   P A 
Sbjct: 2136 QLLVGLLDNPQHRLCELPMLSSAEQ-QVLTGQLQGEHDFDLGQTLHGLFAAQAARTPQAG 2194

Query: 4089 AVKFDAQTLSYGELDRQANRLAHALIARGVGPDVRVAIAMPRSAEIMVAFLAVMKAGGVY 4148
            A+ F  Q L+Y ELD+QANRLA AL  RGVGP VRV +A+ RS E++V  LA++KA G Y
Sbjct: 2195 ALTFAGQHLTYAELDQQANRLARALRERGVGPQVRVGLALERSLEMVVGLLAILKADGAY 2254

Query: 4149 VPLDIEYPRDRLLYMMQDSRAQLLLTHTRALQQL-PIPDGLDSLAI-DRTEEWAGFSDTA 4206
            VPLD EYP DRL YM++DSR  LLL+    L  L  +P+G+ S ++ D     A +SD A
Sbjct: 2255 VPLDPEYPLDRLRYMIEDSRIGLLLSQRALLHTLGELPEGVASWSLEDDAASLAAYSD-A 2313

Query: 4207 PQVKLG-GDNLAYVIYTSGSTGMPKGVAVSHGPLVAHIIATGERYETSPADCELHFMSFA 4265
            P   L    + AY+IYTSGSTG PKGV VSHG    H  A    +     DCELHF S  
Sbjct: 2314 PLDNLSLPQHQAYLIYTSGSTGKPKGVVVSHGEFAMHCQAVIAAFGMRSDDCELHFYSIN 2373

Query: 4266 FDGSHEGWMHPLINGASVLIRDDSLWLPEYTYQQMHRHHVTMAVFPPVYLQQLAEHAERD 4325
            FD + E    PL+ GA V++R    W  E   Q +    V++  F P Y  QLA++    
Sbjct: 2374 FDAASERLWVPLLCGARVVLRAQGQWGAEDICQLVREQQVSILGFTPSYGSQLAQYLGGR 2433

Query: 4326 GNPPKVRVYCFGGDAVAQASYDLAWRALKPTYLFNGYGPTETVVTPLLWKARKGDPCGAV 4385
            G    VR+   GG+A+         +A  P   FN YGPTETVV PL   A +  P    
Sbjct: 2434 GEQLPVRLVITGGEALTGEHLQRIRQAFAPQQCFNAYGPTETVVMPLACLAPEALPADLG 2493

Query: 4386 YAPIGTLLGNRSGYVLDAQLNLQPIGVAGELYLGGEGVARGYLERPALTAERFVPDPFGK 4445
              PIG ++G+R+ Y+LD  L L P G  GELY+GG G+A+GY +RP LTAERFV DPF  
Sbjct: 2494 SVPIGRVIGSRTAYILDEDLALLPQGGIGELYVGGAGLAQGYHDRPGLTAERFVADPFSA 2553

Query: 4446 PGSRVYRSGDLTRGRPDGGVDYLGSVAHQVKIRGFRIELGEIEARLREQDNVGETVVVAH 4505
             G R+YR+GDL R   DG V+Y+G    QVKIRGFRIELGEIE+RL   ++V E VV+A 
Sbjct: 2554 EGGRLYRTGDLVRLGTDGLVEYVGRADQQVKIRGFRIELGEIESRLLAHEDVDEAVVLAL 2613

Query: 4506 EGPTGKQLVAYVVPADTLLADETE--FRDSLRRALKTHLPDYMVPTHFMFLAQMPLTPNG 4563
            + P GKQLV Y+V       ++ +   R++++   + HLPDYMVP H + L  +PL  NG
Sbjct: 2614 DLPGGKQLVGYLVCRQATAGNDVQQLLREAVKADARQHLPDYMVPAHLVLLDSLPLMGNG 2673

Query: 4564 KLDRKGLPEPDASLLQQAYVAPETELEQQIAAIWADVLRLPQVGLNDNFFEVGGHSLLAI 4623
            KLDR  LP PD    +Q Y AP  E+E Q+A IW DVL + +VG+ DNFFE+GG S+L+I
Sbjct: 2674 KLDRHALPLPDLEQARQQYQAPGNEVEAQLAQIWRDVLNVARVGVQDNFFELGGDSILSI 2733

Query: 4624 QITSRVQAELGLEVPLLEVFQTETLRAYV------QAAATFRAG 4661
            Q+ SR + + GL+    ++FQ +T++         +AA+T   G
Sbjct: 2734 QVVSRAR-QAGLQFTPRDLFQHQTIQTLATVVQLSEAASTVEQG 2776



 Score = 1367 bits (3538), Expect = 0.0
 Identities = 981/2593 (37%), Positives = 1359/2593 (52%), Gaps = 160/2593 (6%)

Query: 1545 ADLYPLSPMQQGMLFHTLYEPEAEAYINQLRLDIEG-LDLLAFGRAWQAALDRHDILRSS 1603
            A   P S  Q  +      EP++ AY     L + G LD+ A   A+QA + RH+ LR+ 
Sbjct: 671  AQALPQSLAQNRLWLLWQLEPQSAAYNIPAGLHLRGELDVNALHVAFQALVARHESLRTV 730

Query: 1604 FHWLGLDSAHQVIQRQVDVQLHVIEDIDADCDALAVAERE----KGFTLNAAPLFRLMLV 1659
            F      +  +++ +Q  + LH + D++        A+RE    + F L   PL R+ LV
Sbjct: 731  FSEENGQALQRILPQQ-PLSLHRL-DLEGHSPEQVAAQRETEARQPFDLTQGPLLRVTLV 788

Query: 1660 RGAGTAWHLIFTSHHILMDGWSNAQLLGEVIAHYAGHA-------VPAPLGQFRDYLGWL 1712
            R       L  T HHI+ DGWS   LL E    YA           P PLG + DY  W 
Sbjct: 789  RLGDEEHQLWVTLHHIVADGWSLNILLEEFAKLYAARCQGLEANLAPLPLG-YADYGSWQ 847

Query: 1713 HQQ-ASGEA-----FWKATLA-PLQTPTLLAQALRVPTEGKGMAEHRVLLGDDFTRALGE 1765
             Q  A GEA     +WKA L   L    L     R        A   + +      AL  
Sbjct: 848  RQWLADGEAERQLQYWKAHLGGELPALDLCTDHPRASQREHSAARFSLKVPVRLGEALKG 907

Query: 1766 FARRHKVTLNTVLQGAWSLLLQRYTGQDCVAFGATVAGRSAPLPGIEQQ--LGLFINTLP 1823
             A + + +L  VL   W  LLQRY+GQ  +  G   A R    P +E Q  +G FINT  
Sbjct: 908  LAHQQQASLFMVLLAGWQALLQRYSGQADIRVGVPNANR----PRLETQGMVGFFINTQV 963

Query: 1824 IVSAASPALSVAGWLSELQALNLSLRDHEHVPLYD-IQGWAGQQGAALFDTLLVFENFPV 1882
            + +      S    L++++   L  + ++ +P    ++     +   LF  +   +   +
Sbjct: 964  LRARFDGRHSFTQLLTQVRQATLQAQANQDLPFEQLVEALPDAREQGLFQVMFNHQQRDL 1023

Query: 1883 AEALKQGAPAGLTFGRLHNHEQTHYPLTLGIELGASLRLEFGYDQAR--FSAQQVARLSG 1940
            + AL++  P  L      +  +  + L L  E     RL   +D A   F A  V RL+ 
Sbjct: 1024 S-ALRR-LPGLLAEELAWHSREAKFDLQLHSEEDHLGRLSLAFDYAAELFEASTVERLAH 1081

Query: 1941 NLHHLLVQMLADAQAPLGNLRLLDAATHRDVLAHSRATGAAPRQLRVHERIAAQAAATPD 2000
            +L  LL Q+ A  Q  LG+++LLD      +L   +A  AA + L V E++  Q   TP 
Sbjct: 1082 HLLALLEQVCAAPQQALGDVQLLDEPGRAQLLGWGQAPAAAAQHLLV-EQLNEQVRLTPQ 1140

Query: 2001 ALAVQAGDARLSYAQLNDHANRLAHRLLALGVGPGQRVGLAARRGPQLIVSLLAVLKSGA 2060
              A+      L YA L+  ANRLAH L   GVGP   V +A  R PQL+V LLA+LK+G 
Sbjct: 1141 RTALVWDGGSLDYAGLHQQANRLAHYLRDKGVGPDTCVAIAIERSPQLLVGLLAILKAGG 1200

Query: 2061 AYVPLDPNYPTERLAYMLADSRLDLLLSETGLLADLPLPQGLARLDFTASGAELADYPST 2120
            AYVPLD +YP ERLAYMLAD    LLLS + LL  LP   G++ +        L  +PS 
Sbjct: 1201 AYVPLDVDYPAERLAYMLADCDARLLLSHSSLLGKLPQVDGVSAI--ALDQLHLDSWPSH 1258

Query: 2121 PPANHAAAADLAYVIYTSGSTGLPKGVAIDHAALSQFCDSAEAYSRLSASDRVLQFATFS 2180
             P  H    +LAYVIYTSGSTG PKGV   HAAL++     +    L+ SD ++Q A  S
Sbjct: 1259 APGLHLHGDNLAYVIYTSGSTGQPKGVGNTHAALAERLQWMQTTYALNDSDVLMQKAPIS 1318

Query: 2181 FDGFVEQCYPPLCVGAALIMRGD-ELWDAGQLAREIVEQGVTLADLPAAYWYLLAKECAV 2239
            FD  V +C+ PL  G  L++ G  E  D  ++A  +   GVT          +  +E   
Sbjct: 1319 FDVSVWECFWPLVTGCKLVLAGPGEHRDPQRIAALVQAYGVTTLHFVPPLLQVFVQEPQA 1378

Query: 2240 DRRTLGNLRQVHVGGEAMSVEGLRAWHTAGLGGVRLVNTYGPTEATVVSSVHDCQLADAS 2299
                  +LR V  GGEA+S   LR      L  V+L N YGPTE  +  +   CQ+ D  
Sbjct: 1379 S--ACSSLRWVFSGGEALSAT-LRDRVLQVLPQVQLHNRYGPTETAINVTHWHCQVVDGE 1435

Query: 2300 DAFGVPIGRAIEGRALYVLDSGFELLASDGVGELCIGAEHGLAQCYFDRPALTAERFLPD 2359
             +   PIGR +      VLD   EL A    GEL +G   GLA+ Y  RP LTAERF+P 
Sbjct: 1436 RS---PIGRPLGNVLCRVLDDELELSAPGVPGELYLGGA-GLARGYLGRPGLTAERFVPQ 1491

Query: 2360 PFSAVPGARLYRSGDLARYN-EAGALEYVGRIDHQVKIRGFRVEMGEIEASLQALPSLRE 2418
                  G RLYRSGD AR+  +  ALEY+GR+D QVK+RGFRVE  E++A L A   + +
Sbjct: 1492 ADGN--GQRLYRSGDRARWQVQLEALEYLGRLDQQVKVRGFRVEPEEVQACLLAQAGVEQ 1549

Query: 2419 AAVIAQPSATGTQLVAYVVPVTAQTLDPQALAATLRQSLPDYMVPGQWVILEALPLNNNG 2478
            A V+    A G QLV Y     + +  P  + A L + LP YMVP Q + L  +PL  +G
Sbjct: 1550 ALVLIHKDAIGAQLVGYY----SGSAQPAEVLAVLAEQLPAYMVPAQLIPLAQMPLGPSG 1605

Query: 2479 KLDRRALPAPDLNQARQAYLAPQNPLQSRLAAIWQAVLQVEQVGLNDHFFERGGHSLLAT 2538
            K+DR+ALPAP   Q  + ++ P+  LQ ++AAIW+ VL + ++GL D FF  GGHSLLAT
Sbjct: 1606 KVDRKALPAPVWQQ--REHVEPRTELQQQVAAIWRDVLNLPRIGLQDDFFALGGHSLLAT 1663

Query: 2539 QVVSRVRHDLKLEVPLRALFEQPTLEAFAAACAGLQTAPALPL----VAVTRDQPLALSF 2594
            Q+VSR R    +E+PL+ALFE   L AF A  A L+ A    L      V R Q + LS+
Sbjct: 1664 QIVSRTRQACDVELPLKALFEASELGAFCAEIARLRAAGERNLQGAITRVDRRQAVPLSY 1723

Query: 2595 AQERQWFLWQLDPTSAAYHVPSALHLRGRLDVAALEQAFQVLVQRHEPLRTTFVEEGEHT 2654
            +Q+R WFLWQ++P S AY+V     LRG L V A E+A Q L+ RHE LRTTF       
Sbjct: 1724 SQQRMWFLWQMEPDSPAYNVGGMARLRGTLHVDAFERALQALIVRHETLRTTFPSIDGVP 1783

Query: 2655 WQVIHPRLAMPIERQRVVASAIEHAVAQEIQR--------PFDLVNGPLMRVNLLEVEAE 2706
            +Q +     + ++ Q   A + + A  Q +Q+        PFDL  GPL+R  L++    
Sbjct: 1784 YQCVAEDSGLHLDWQDFSALS-DDACQQRLQQLADDQAHQPFDLERGPLLRACLVKANER 1842

Query: 2707 HHVLVITQHHIISDGWSMQVMVDELVALYQG-----QAELPALPIQYVDYAQWQRDWMAA 2761
             H  V+T HHI+++GW+M +   EL  LY+      ++ L  LP+QY+DY+ WQR W+ +
Sbjct: 1843 EHFFVLTLHHIVTEGWAMDIFARELGELYEAFVDDRESPLAPLPVQYLDYSVWQRQWLES 1902

Query: 2762 GEKQRQLDYWRTRLGTEHPVLELPLDHPRPALQSHRGARRQVQLDNALVAELKALAQRQE 2821
            GE  RQL YW+ RLG EHPVL LP D PRPA+QSHRG   + +LD ALVA + A   ++ 
Sbjct: 1903 GEGARQLAYWKHRLGDEHPVLALPADRPRPAVQSHRGELYRFELDPALVARVHAFNSQRG 1962

Query: 2822 VTLFVLLLASFQTLLHRYSGQSRIRVGVPIANRNRLETERLLGFFVNTQVLQADVHGQMA 2881
            +TLF+ + A+   LLHRYSGQ  +R+G P+ANR R E+E L+G F+NTQVL+ ++ GQM 
Sbjct: 1963 LTLFMTMTATLAALLHRYSGQRDLRIGAPVANRIRPESEGLIGAFLNTQVLRCELDGQMT 2022

Query: 2882 FDQLLAQVKQQALDAQAHQDLPFEQLVEALQPERSLSHNPLFQVMFNHQDNLRAAPLQLP 2941
               LL QV+Q A++ Q+HQDLPF+QLVEALQP RS ++NPLFQVM N Q         L 
Sbjct: 2023 ASALLEQVRQAAIEGQSHQDLPFDQLVEALQPPRSSAYNPLFQVMCNVQRWAFQQSRTLA 2082

Query: 2942 GLDLQAVDWAGHSTQFDLNLETEESAEGLWASLTYATDLFDATTVERLAEHWQNLLRAVV 3001
            G+ +  +     +T+FDL LE  +    L   LTY+ DLFDA  + R+AEHWQ LL  ++
Sbjct: 2083 GMQVDYLVNDASATKFDLYLEVTDLDGRLGCCLTYSRDLFDAPRIARMAEHWQQLLVGLL 2142

Query: 3002 NDASQAVDELAMLSAPQWQQMVEAWNDTSVDYPRERCVHTLFEARALAQPDALALQFKGQ 3061
            ++    + EL MLS+ + QQ++        D+   + +H LF A+A   P A AL F GQ
Sbjct: 2143 DNPQHRLCELPMLSSAE-QQVLTGQLQGEHDFDLGQTLHGLFAAQAARTPQAGALTFAGQ 2201

Query: 3062 ALSYGELNRRANRLAHRLIAAGVGPDVLVAVHVERSLDMVIGLLATLKAGGAYVPLDPQF 3121
             L+Y EL+++ANRLA  L   GVGP V V + +ERSL+MV+GLLA LKA GAYVPLDP++
Sbjct: 2202 HLTYAELDQQANRLARALRERGVGPQVRVGLALERSLEMVVGLLAILKADGAYVPLDPEY 2261

Query: 3122 PADRLAFMLEDSRARVLLTQSHLDGSLAH-PQGVQVLMVEADDN---AVEHDPQVNVT-P 3176
            P DRL +M+EDSR  +LL+Q  L  +L   P+GV    +E D     A    P  N++ P
Sbjct: 2262 PLDRLRYMIEDSRIGLLLSQRALLHTLGELPEGVASWSLEDDAASLAAYSDAPLDNLSLP 2321

Query: 3177 EHLAYVIYTSGSTGTPKGVMVRHEALCSFTCGMAGTLDIGADARVLSLTTFSFDIFALEL 3236
            +H AY+IYTSGSTG PKGV+V H         +     + +D   L   + +FD  +  L
Sbjct: 2322 QHQAYLIYTSGSTGKPKGVVVSHGEFAMHCQAVIAAFGMRSDDCELHFYSINFDAASERL 2381

Query: 3237 YVPLTVGATVLLSGQELALDPEAIIDLVHSQCANVLQATPSTWRMLLE--SPRAQLLRGI 3294
            +VPL  GA V+L  Q      E I  LV  Q  ++L  TPS    L +    R + L   
Sbjct: 2382 WVPLLCGARVVLRAQG-QWGAEDICQLVREQQVSILGFTPSYGSQLAQYLGGRGEQLPVR 2440

Query: 3295 KCLCGGEALPSDLAQRLLDLQGP--LWNLYGPTETTIWSAAHRLHEAQPL------VGRP 3346
              + GGEAL  +  QR+     P   +N YGPTET +   A    EA P       +GR 
Sbjct: 2441 LVITGGEALTGEHLQRIRQAFAPQQCFNAYGPTETVVMPLACLAPEALPADLGSVPIGRV 2500

Query: 3347 IANTALFILNAGLTPSPVGAAGELLIGGVGLARGYHERPALTAERFVPNPYGAPGERLYR 3406
            I +   +IL+  L   P G  GEL +GG GLA+GYH+RP LTAERFV +P+ A G RLYR
Sbjct: 2501 IGSRTAYILDEDLALLPQGGIGELYVGGAGLAQGYHDRPGLTAERFVADPFSAEGGRLYR 2560

Query: 3407 TGDLARYRADGVVEYIGRVDHQVKVRGFRIELGEVEACLREQAGIREAVVLA----DNDR 3462
            TGDL R   DG+VEY+GR D QVK+RGFRIELGE+E+ L     + EAVVLA       +
Sbjct: 2561 TGDLVRLGTDGLVEYVGRADQQVKIRGFRIELGEIESRLLAHEDVDEAVVLALDLPGGKQ 2620

Query: 3463 LIAYLVSSAPDAP--------EVYKAALRERLPDYMVPAHLMFLDSLPLTPNGKLDRKAL 3514
            L+ YLV     A         E  KA  R+ LPDYMVPAHL+ LDSLPL  NGKLDR AL
Sbjct: 2621 LVGYLVCRQATAGNDVQQLLREAVKADARQHLPDYMVPAHLVLLDSLPLMGNGKLDRHAL 2680

Query: 3515 PAVDAALLRTGHVAPVTPREQQVAAIWADVLDLPQVGLDDHFFELGGHSLLATRVVSRVR 3574
            P  D    R  + AP    E Q+A IW DVL++ +VG+ D+FFELGG S+L+ +VVSR R
Sbjct: 2681 PLPDLEQARQQYQAPGNEVEAQLAQIWRDVLNVARVGVQDNFFELGGDSILSIQVVSRAR 2740

Query: 3575 QALALEVALKTLFEHPLLGDF--VQALGEEGVVAPALLKADRRQPLPLSYAQERQWFLWQ 3632
            QA  L+   + LF+H  +     V  L E        L+  +    P+ +     WF   
Sbjct: 2741 QA-GLQFTPRDLFQHQTIQTLATVVQLSEAASTVEQGLRQGQAGLTPIQH-----WFFDS 2794

Query: 3633 LDPHSTAYHIPSALRLHGQLDLGALQRSFDTLLARHESLRTHVRQDA-----AATVQVIE 3687
              P    ++    L +   L    L+++   L+  H+SLR     DA     A  VQ   
Sbjct: 2795 DVPQPQHWNQTVLLEVRQPLVEATLEQALAALVQHHDSLRLRF-SDAQGRWQAQYVQPGA 2853

Query: 3688 DSGLIEIDLADTDEASLKARVAEVVAQPFELLRGPLLRVKLLRLAADDHVLVLVQHHIVS 3747
            +  L    +AD ++      +   V +  +L +GPLLR  L+R       L+L  HH+V 
Sbjct: 2854 EQLLWSATVADFNDCQA---LYTDVQRSLDLQQGPLLRALLVRDGQGTQRLLLAIHHLVV 2910

Query: 3748 DGWSMQLMVEELVQLYAAFSQGHMPDLPALPIQYADYALWQRNWMEAGEKARQLAYWQAQ 3807
            DG S ++++E+L  LY          LPA      D+A    ++  +     +LA+WQ Q
Sbjct: 2911 DGVSWRVLLEDLQALYRG------QPLPAKTHAVGDWAARLASYAGSDSLRDELAWWQGQ 2964

Query: 3808 LGGVQPVLELPFDYLRPAVQSHRGARLGIELQPQLLAGLRRLAQGAGVT-LPMVLLASYQ 3866
            LGGV+   ELP D+ + +        L I L  +    L +LA  A  T +  +LL +  
Sbjct: 2965 LGGVR--RELPCDHPQGSNLHRHAQTLAIGLDVEQTRQLLQLAPAAYHTQVNDLLLTALA 3022

Query: 3867 ALLHRYSGQEDVRVGVPIANRNRL----ETEGLIGFFVNTQVL-------KADIHGQMSV 3915
              L R+SG E+V V +    R+ L    +    +G+F N   L       + D     S+
Sbjct: 3023 RALCRWSGDEEVLVQLEGHGRDGLFEDMDLTRSVGWFTNAYPLSLRPLPGEDDAARAGSI 3082

Query: 3916 VQLLQQVRQRSLEAQAHQDLPFEQLVEALQPERSMSLSPLFQVLFNH-----RVTSAAHH 3970
             ++ +Q+R    +   +  L +  L +A   E+ M+  P  ++ FN+     +   +A  
Sbjct: 3083 KRIKEQLRHVPHKGLGYGVLRY--LADAAGREQ-MAALPQARITFNYLGQFDQQFDSAAL 3139

Query: 3971 LQRLT-------ELDVEVLSWDEGVAQFDLALDVEESQATLRASLNYATDLFAPATIERM 4023
             Q L        +LD  + +W        L++D +     L+    ++ + +   TI R+
Sbjct: 3140 FQPLEASAGLAHDLDAPLPNW--------LSVDGQVYGGALQLRWTFSAERYDQQTIARL 3191

Query: 4024 AGHWQNLLQAMVA 4036
            A  ++  L A+VA
Sbjct: 3192 AEGYRQELLALVA 3204



 Score =  967 bits (2500), Expect = 0.0
 Identities = 611/1513 (40%), Positives = 844/1513 (55%), Gaps = 84/1513 (5%)

Query: 46   VEDRQAL--SYAQQRMWFLWQLDPASGAYNLPGAVRLKGALSLDALEQAFASLVARHETL 103
            +E  QAL  S AQ R+W LWQL+P S AYN+P  + L+G L ++AL  AF +LVARHE+L
Sbjct: 668  LERAQALPQSLAQNRLWLLWQLEPQSAAYNIPAGLHLRGELDVNALHVAFQALVARHESL 727

Query: 104  RTVFQRQADERLLQVAADTSLTVEQLDLSHLALAEREEAVNAAATRQSLLPFDLENGPLL 163
            RTVF  +  + L ++     L++ +LDL         E V A    ++  PFDL  GPLL
Sbjct: 728  RTVFSEENGQALQRILPQQPLSLHRLDLE----GHSPEQVAAQRETEARQPFDLTQGPLL 783

Query: 164  RVQLLKLDAQEHVLLLTLHHIVSDGWSMNVLIDEFIRCYDAHERSQAPQLPALPIQYSDY 223
            RV L++L  +EH L +TLHHIV+DGWS+N+L++EF + Y A  +     L  LP+ Y+DY
Sbjct: 784  RVTLVRLGDEEHQLWVTLHHIVADGWSLNILLEEFAKLYAARCQGLEANLAPLPLGYADY 843

Query: 224  ALWQRRWLEAGEQARQLEYWQARLGDEHPVLELPTDRPRPAMPSYQGTRHDFAINPQLAA 283
              WQR+WL  GE  RQL+YW+A LG E P L+L TD PR +   +   R    +  +L  
Sbjct: 844  GSWQRQWLADGEAERQLQYWKAHLGGELPALDLCTDHPRASQREHSAARFSLKVPVRLGE 903

Query: 284  QLRTCAQKHNVTLFMLLLGAFNVLLHRYTGQGDIRVGVPIANRNRTEVEGLIGFFVNTQV 343
             L+  A +   +LFM+LL  +  LL RY+GQ DIRVGVP ANR R E +G++GFF+NTQV
Sbjct: 904  ALKGLAHQQQASLFMVLLAGWQALLQRYSGQADIRVGVPNANRPRLETQGMVGFFINTQV 963

Query: 344  LRTELTGQTLVTELLQSIKEHALGAQAHQELPFERLVEALKVERSLSHTPLFQVMYNHQP 403
            LR    G+   T+LL  +++  L AQA+Q+LPFE+LVEAL   R      LFQVM+NHQ 
Sbjct: 964  LRARFDGRHSFTQLLTQVRQATLQAQANQDLPFEQLVEALPDAREQG---LFQVMFNHQQ 1020

Query: 404  VVADIANVSTASGLELALVEWQGRTTQFDLTLDTYEKS-GTLHAALTYANDLFDALTIER 462
               D++ +    GL    + W  R  +FDL L + E   G L  A  YA +LF+A T+ER
Sbjct: 1021 --RDLSALRRLPGLLAEELAWHSREAKFDLQLHSEEDHLGRLSLAFDYAAELFEASTVER 1078

Query: 463  MAGHWIRLLQAMVADGTQRIGELPMLAADEQQLLVHAWNQTAEAYPTERGIHHLVE---D 519
            +A H + LL+ + A   Q +G++ +L    +  L+  W Q     P     H LVE   +
Sbjct: 1079 LAHHLLALLEQVCAAPQQALGDVQLLDEPGRAQLL-GWGQA----PAAAAQHLLVEQLNE 1133

Query: 520  QVQATPDAPALVFGTTTLTYAQLDARANQLAHALREKGVAPDVLVGICIERSIEMVVGLL 579
            QV+ TP   ALV+   +L YA L  +AN+LAH LR+KGV PD  V I IERS +++VGLL
Sbjct: 1134 QVRLTPQRTALVWDGGSLDYAGLHQQANRLAHYLRDKGVGPDTCVAIAIERSPQLLVGLL 1193

Query: 580  AILKAGGAYVPLDPEYPQERLAYMIEDSGIQLLLSQHSLLPVLP-VEGIQVIALDQPASW 638
            AILKAGGAYVPLD +YP ERLAYM+ D   +LLLS  SLL  LP V+G+  IALDQ    
Sbjct: 1194 AILKAGGAYVPLDVDYPAERLAYMLADCDARLLLSHSSLLGKLPQVDGVSAIALDQ--LH 1251

Query: 639  LDGYSAESANVSLHALNLAYVIYTSGSTGKPKGAGNSHRALVNRLCWMQQAYGLDARDAV 698
            LD + + +  + LH  NLAYVIYTSGSTG+PKG GN+H AL  RL WMQ  Y L+  D +
Sbjct: 1252 LDSWPSHAPGLHLHGDNLAYVIYTSGSTGQPKGVGNTHAALAERLQWMQTTYALNDSDVL 1311

Query: 699  LQKTPFSFDVSVWEFFWPLMTGARLVVAAPGEHREPARLIETIGRHGITTLHFVPSMLQA 758
            +QK P SFDVSVWE FWPL+TG +LV+A PGEHR+P R+   +  +G+TTLHFVP +LQ 
Sbjct: 1312 MQKAPISFDVSVWECFWPLVTGCKLVLAGPGEHRDPQRIAALVQAYGVTTLHFVPPLLQV 1371

Query: 759  FIHEPGVQACTSLQRIVCSGEALPLDAQLQVFAKLPQAGLFNLYGPTEAAIDVTHWTCVD 818
            F+ EP   AC+SL+ +   GEAL    + +V   LPQ  L N YGPTE AI+VTHW C  
Sbjct: 1372 FVQEPQASACSSLRWVFSGGEALSATLRDRVLQVLPQVQLHNRYGPTETAINVTHWHCQV 1431

Query: 819  EGADSVPIGRPIANLGTYVLDAQLNPVPAGVSGELYLGGIGLARSYHRRPALTAERFVPS 878
               +  PIGRP+ N+   VLD +L     GV GELYLGG GLAR Y  RP LTAERFVP 
Sbjct: 1432 VDGERSPIGRPLGNVLCRVLDDELELSAPGVPGELYLGGAGLARGYLGRPGLTAERFVPQ 1491

Query: 879  PFGDGARLYRTGDRVRQRAD-GVIEYLGRLDHQVKLRGLRIELGEIETRLMQHPSVREAV 937
              G+G RLYR+GDR R +     +EYLGRLD QVK+RG R+E  E++  L+    V +A+
Sbjct: 1492 ADGNGQRLYRSGDRARWQVQLEALEYLGRLDQQVKVRGFRVEPEEVQACLLAQAGVEQAL 1551

Query: 938  VLVQG---GKQLVAYLVLEDTAPVDLKAWLLSSLPEYMVPSHMVPLAKLPVTANGKLDRK 994
            VL+     G QLV Y       P ++ A L   LP YMVP+ ++PLA++P+  +GK+DRK
Sbjct: 1552 VLIHKDAIGAQLVGY-YSGSAQPAEVLAVLAEQLPAYMVPAQLIPLAQMPLGPSGKVDRK 1610

Query: 995  ALPLPDVAPQQAFVAPENDLQKAVAAIWSDVLGVEQVGLDDNFFELGGDSIISIQVVSRA 1054
            ALP P V  Q+  V P  +LQ+ VAAIW DVL + ++GL D+FF LGG S+++ Q+VSR 
Sbjct: 1611 ALPAP-VWQQREHVEPRTELQQQVAAIWRDVLNLPRIGLQDDFFALGGHSLLATQIVSRT 1669

Query: 1055 RQA-GIRLSPRDLFQYQSVRSLALVATFEQATAIDQGPVTGEVI------LTPVQHS--- 1104
            RQA  + L  + LF+   +   A  A   +  A  +  + G +         P+ +S   
Sbjct: 1670 RQACDVELPLKALFEASELG--AFCAEIARLRAAGERNLQGAITRVDRRQAVPLSYSQQR 1727

Query: 1105 --FFEQAIPARQHWNQSLLLTPREALEPARLEAALTRLINHHDALRLRFLQQADGWRQAH 1162
              F  Q  P    +N   +   R  L     E AL  LI  H+ LR  F        Q  
Sbjct: 1728 MWFLWQMEPDSPAYNVGGMARLRGTLHVDAFERALQALIVRHETLRTTFPSIDGVPYQCV 1787

Query: 1163 AEPVATASLWQ--SQVADEA---ELAALC-DEAQRSLDLEQGPLLRAVLAVMADGTQRLL 1216
            AE       WQ  S ++D+A    L  L  D+A +  DLE+GPLLRA L    +     +
Sbjct: 1788 AEDSGLHLDWQDFSALSDDACQQRLQQLADDQAHQPFDLERGPLLRACLVKANEREHFFV 1847

Query: 1217 LVVHHLAVDGVSWRILLEDLQQAYRN---------AALPAKTSAYQLWAQQLQRHAQTLD 1267
            L +HH+  +G +  I   +L + Y           A LP +   Y +W +Q     +   
Sbjct: 1848 LTLHHIVTEGWAMDIFARELGELYEAFVDDRESPLAPLPVQYLDYSVWQRQWLESGEGA- 1906

Query: 1268 EQLPYWQAQTASAD----LPCDNPQGGLQNRLGSLLEIRLSAEHTRQLLQDAPAAYRTQV 1323
             QL YW+ +         LP D P+  +Q+  G L    L      ++   A  + R   
Sbjct: 1907 RQLAYWKHRLGDEHPVLALPADRPRPAVQSHRGELYRFELDPALVARV--HAFNSQRGLT 1964

Query: 1324 NDLLLTA-LARVISRWSEQPAALIQLEGHGREDLFDSVDLSRTVGWFTSLFPVRLHADGE 1382
              + +TA LA ++ R+S Q    I      R       +    +G F +   +R   DG+
Sbjct: 1965 LFMTMTATLAALLHRYSGQRDLRIGAPVANR----IRPESEGLIGAFLNTQVLRCELDGQ 2020

Query: 1383 LSAAIKSVKEQLRAVPHKGIGYGLLRYLGTPDAREALSALAAPRIT-----FNYLGQFDR 1437
            ++A+  ++ EQ+R    +G  +  L +       + + AL  PR +     F  +    R
Sbjct: 2021 MTAS--ALLEQVRQAAIEGQSHQDLPF------DQLVEALQPPRSSAYNPLFQVMCNVQR 2072

Query: 1438 QFNESALFVPATQGSGQAQDPQAPLANWLTVEGQVYGGELALQWGFSREMFDVATVQRLA 1497
               + +  +   Q      D  A   + L +E     G L     +SR++FD   + R+A
Sbjct: 2073 WAFQQSRTLAGMQVDYLVNDASATKFD-LYLEVTDLDGRLGCCLTYSRDLFDAPRIARMA 2131

Query: 1498 DDYRAELQALIEH 1510
            + ++  L  L+++
Sbjct: 2132 EHWQQLLVGLLDN 2144



 Score =  608 bits (1567), Expect = e-176
 Identities = 528/1581 (33%), Positives = 750/1581 (47%), Gaps = 95/1581 (6%)

Query: 1482 GFSREMFDVATVQRLADDYRAELQALIEHCCATPAGQVTPSDFPLARLTQQQLDGLPIAG 1541
            G S     + +  R A D    L+AL E   A+  G        L    ++ L G  I  
Sbjct: 1657 GHSLLATQIVSRTRQACDVELPLKALFE---ASELGAFCAEIARLRAAGERNLQGA-ITR 1712

Query: 1542 PAIADLYPLSPMQQGMLFHTLYEPEAEAYINQLRLDIEG-LDLLAFGRAWQAALDRHDIL 1600
                   PLS  QQ M F    EP++ AY       + G L + AF RA QA + RH+ L
Sbjct: 1713 VDRRQAVPLSYSQQRMWFLWQMEPDSPAYNVGGMARLRGTLHVDAFERALQALIVRHETL 1772

Query: 1601 RSSFHWLGLDSA-HQVIQRQVDVQLHVI-EDIDADCD--------ALAVAEREKGFTLNA 1650
            R++F    +D   +Q +    D  LH+  +D  A  D         LA  +  + F L  
Sbjct: 1773 RTTFP--SIDGVPYQCVAE--DSGLHLDWQDFSALSDDACQQRLQQLADDQAHQPFDLER 1828

Query: 1651 APLFRLMLVRGAGTAWHLIFTSHHILMDGWSN---AQLLGEVIAHYAGHA----VPAPLG 1703
             PL R  LV+        + T HHI+ +GW+    A+ LGE+   +         P P+ 
Sbjct: 1829 GPLLRACLVKANEREHFFVLTLHHIVTEGWAMDIFARELGELYEAFVDDRESPLAPLPV- 1887

Query: 1704 QFRDYLGWLHQQA-SGE-----AFWKATLAPLQTPTLLAQALRV-PTEGKGMAE-HRVLL 1755
            Q+ DY  W  Q   SGE     A+WK  L   + P L   A R  P       E +R  L
Sbjct: 1888 QYLDYSVWQRQWLESGEGARQLAYWKHRLGD-EHPVLALPADRPRPAVQSHRGELYRFEL 1946

Query: 1756 GDDFTRALGEFARRHKVTLNTVLQGAWSLLLQRYTGQDCVAFGATVAGRSAPLPGIEQQL 1815
                   +  F  +  +TL   +    + LL RY+GQ  +  GA VA R  P    E  +
Sbjct: 1947 DPALVARVHAFNSQRGLTLFMTMTATLAALLHRYSGQRDLRIGAPVANRIRPES--EGLI 2004

Query: 1816 GLFINTLPIVSAASPALSVAGWLSELQALNLSLRDHEHVPLYD-IQGWAGQQGAALFDTL 1874
            G F+NT  +       ++ +  L +++   +  + H+ +P    ++     + +A     
Sbjct: 2005 GAFLNTQVLRCELDGQMTASALLEQVRQAAIEGQSHQDLPFDQLVEALQPPRSSAYNPLF 2064

Query: 1875 LVFENFPVAEALKQGAPAGLTFGRLHNHEQ-THYPLTLGI-ELGASLRLEFGYDQARFSA 1932
             V  N       +    AG+    L N    T + L L + +L   L     Y +  F A
Sbjct: 2065 QVMCNVQRWAFQQSRTLAGMQVDYLVNDASATKFDLYLEVTDLDGRLGCCLTYSRDLFDA 2124

Query: 1933 QQVARLSGNLHHLLVQMLADAQAPLGNLRLLDAATHRDVLAHSRATGAAPRQLRVHERIA 1992
             ++AR++ +   LLV +L + Q  L  L +L +A  + +    +          +H   A
Sbjct: 2125 PRIARMAEHWQQLLVGLLDNPQHRLCELPMLSSAEQQVLTGQLQGEHDFDLGQTLHGLFA 2184

Query: 1993 AQAAATPDALAVQAGDARLSYAQLNDHANRLAHRLLALGVGPGQRVGLAARRGPQLIVSL 2052
            AQAA TP A A+      L+YA+L+  ANRLA  L   GVGP  RVGLA  R  +++V L
Sbjct: 2185 AQAARTPQAGALTFAGQHLTYAELDQQANRLARALRERGVGPQVRVGLALERSLEMVVGL 2244

Query: 2053 LAVLKSGAAYVPLDPNYPTERLAYMLADSRLDLLLSETGLLADL-PLPQGLARLDFTASG 2111
            LA+LK+  AYVPLDP YP +RL YM+ DSR+ LLLS+  LL  L  LP+G+A        
Sbjct: 2245 LAILKADGAYVPLDPEYPLDRLRYMIEDSRIGLLLSQRALLHTLGELPEGVASWSLEDDA 2304

Query: 2112 AELADYPSTPPANHAAAADLAYVIYTSGSTGLPKGVAIDHAALSQFCDSAEAYSRLSASD 2171
            A LA Y   P  N +     AY+IYTSGSTG PKGV + H   +  C +  A   + + D
Sbjct: 2305 ASLAAYSDAPLDNLSLPQHQAYLIYTSGSTGKPKGVVVSHGEFAMHCQAVIAAFGMRSDD 2364

Query: 2172 RVLQFATFSFDGFVEQCYPPLCVGAALIMRGDELWDAGQLAREIVEQGVTLADLPAAYWY 2231
              L F + +FD   E+ + PL  GA +++R    W A  + + + EQ V++     +Y  
Sbjct: 2365 CELHFYSINFDAASERLWVPLLCGARVVLRAQGQWGAEDICQLVREQQVSILGFTPSYGS 2424

Query: 2232 LLAKECAVDRRTLGNLRQVHVGGEAMSVEGLRAWHTAGLGGVRLVNTYGPTEATVVSSVH 2291
             LA+        L  +R V  GGEA++ E L+    A     +  N YGPTE  V+    
Sbjct: 2425 QLAQYLGGRGEQLP-VRLVITGGEALTGEHLQRIRQA-FAPQQCFNAYGPTETVVMPLAC 2482

Query: 2292 DCQLADASDAFGVPIGRAIEGRALYVLDSGFELLASDGVGELCIGAEHGLAQCYFDRPAL 2351
                A  +D   VPIGR I  R  Y+LD    LL   G+GEL +G   GLAQ Y DRP L
Sbjct: 2483 LAPEALPADLGSVPIGRVIGSRTAYILDEDLALLPQGGIGELYVGGA-GLAQGYHDRPGL 2541

Query: 2352 TAERFLPDPFSAVPGARLYRSGDLARYNEAGALEYVGRIDHQVKIRGFRVEMGEIEASLQ 2411
            TAERF+ DPFSA  G RLYR+GDL R    G +EYVGR D QVKIRGFR+E+GEIE+ L 
Sbjct: 2542 TAERFVADPFSA-EGGRLYRTGDLVRLGTDGLVEYVGRADQQVKIRGFRIELGEIESRLL 2600

Query: 2412 ALPSLREAAVIAQPSATGTQLVAYVVPVTA------QTLDPQALAATLRQSLPDYMVPGQ 2465
            A   + EA V+A     G QLV Y+V   A      Q L  +A+ A  RQ LPDYMVP  
Sbjct: 2601 AHEDVDEAVVLALDLPGGKQLVGYLVCRQATAGNDVQQLLREAVKADARQHLPDYMVPAH 2660

Query: 2466 WVILEALPLNNNGKLDRRALPAPDLNQARQAYLAPQNPLQSRLAAIWQAVLQVEQVGLND 2525
             V+L++LPL  NGKLDR ALP PDL QARQ Y AP N ++++LA IW+ VL V +VG+ D
Sbjct: 2661 LVLLDSLPLMGNGKLDRHALPLPDLEQARQQYQAPGNEVEAQLAQIWRDVLNVARVGVQD 2720

Query: 2526 HFFERGGHSLLATQVVSRVRHDLKLEVPLRALFEQPTLEAFAAACAGLQTAPALPLVAVT 2585
            +FFE GG S+L+ QVVSR R    L+   R LF+  T++  A       T   L   A T
Sbjct: 2721 NFFELGGDSILSIQVVSRARQ-AGLQFTPRDLFQHQTIQTLA-------TVVQLSEAAST 2772

Query: 2586 RDQPLALSFA---QERQWFLWQLDPTSAAYHVPSALHLRGRLDVAALEQAFQVLVQRHEP 2642
             +Q L    A     + WF     P    ++    L +R  L  A LEQA   LVQ H+ 
Sbjct: 2773 VEQGLRQGQAGLTPIQHWFFDSDVPQPQHWNQTVLLEVRQPLVEATLEQALAALVQHHDS 2832

Query: 2643 LRTTFVE-EGEHTWQVIHPRLAMPIERQRVVASAIEHAVAQEIQRPFDLVNGPLMRVNLL 2701
            LR  F + +G    Q + P     +    V       A+  ++QR  DL  GPL+R  L+
Sbjct: 2833 LRLRFSDAQGRWQAQYVQPGAEQLLWSATVADFNDCQALYTDVQRSLDLQQGPLLRALLV 2892

Query: 2702 EVEAEHHVLVITQHHIISDGWSMQVMVDELVALYQGQAELPALPIQYVDYAQWQRDWMAA 2761
                    L++  HH++ DG S +V++++L ALY+GQ  LPA      D+A     +  +
Sbjct: 2893 RDGQGTQRLLLAIHHLVVDGVSWRVLLEDLQALYRGQ-PLPAKTHAVGDWAARLASYAGS 2951

Query: 2762 GEKQRQLDYWRTRLGTEHPVLELPLDHPRPALQSHRGARRQVQLDNALVAELKALAQRQE 2821
               + +L +W+ +LG      ELP DHP+ +          + LD     +L  LA    
Sbjct: 2952 DSLRDELAWWQGQLGGVR--RELPCDHPQGSNLHRHAQTLAIGLDVEQTRQLLQLAPAAY 3009

Query: 2822 VT-LFVLLLASFQTLLHRYSGQSRIRVGVPIANRNRL----ETERLLGFFVNTQVLQ--- 2873
             T +  LLL +    L R+SG   + V +    R+ L    +  R +G+F N   L    
Sbjct: 3010 HTQVNDLLLTALARALCRWSGDEEVLVQLEGHGRDGLFEDMDLTRSVGWFTNAYPLSLRP 3069

Query: 2874 ADVHGQMAFDQLLAQVKQQALDAQAHQDLPF---EQLVEALQPERSLSHNPLFQVMFNH- 2929
                   A    + ++K+Q L    H+ L +     L +A   E+ ++  P  ++ FN+ 
Sbjct: 3070 LPGEDDAARAGSIKRIKEQ-LRHVPHKGLGYGVLRYLADAAGREQ-MAALPQARITFNYL 3127

Query: 2930 ---QDNLRAAPLQLPGLDLQAVDWAGHSTQFDLNLETEESAEG------LWASLTYATDL 2980
                    +A L  P   L+A   AG +   D  L    S +G      L    T++ + 
Sbjct: 3128 GQFDQQFDSAALFQP---LEAS--AGLAHDLDAPLPNWLSVDGQVYGGALQLRWTFSAER 3182

Query: 2981 FDATTVERLAEHWQNLLRAVV 3001
            +D  T+ RLAE ++  L A+V
Sbjct: 3183 YDQQTIARLAEGYRQELLALV 3203



 Score =  502 bits (1293), Expect = e-145
 Identities = 375/1082 (34%), Positives = 550/1082 (50%), Gaps = 78/1082 (7%)

Query: 2578 ALPLVAVTRDQPLALSFAQERQWFLWQLDPTSAAYHVPSALHLRGRLDVAALEQAFQVLV 2637
            ALP+ A   +    L+  QE       L+P +  Y++     +   LD     QA+Q +V
Sbjct: 3231 ALPVPAAQIEDVYPLTPMQEGMLLHTLLEPGTGLYYMQDRYRINSALDPERFAQAWQAVV 3290

Query: 2638 QRHEPLRTTFV-EEGEHTWQVIHP--RLAMPIERQRVVAS-----AIEHAVAQEIQRPFD 2689
             RHE LR +F    GE   Q+IH    LA+  +  R +A       ++    QE +  F 
Sbjct: 3291 ARHEALRASFSWNSGEAMLQIIHKPGNLAVDYQDWRGLADDAQEQRLQALHKQEREAGFA 3350

Query: 2690 LVNGPLMRVNLLEVEAEHHVLVITQHHIISDGWSMQVMVDELVALYQG-----QAELPAL 2744
            L++     + L+ V  E +  +++ HHI+ D W   +++++   +YQ      QA+LP +
Sbjct: 3351 LLSEAPFHLRLVRVAEERYWFMMSNHHILIDAWCRSLLMNDFFEVYQALGEGRQAQLP-V 3409

Query: 2745 PIQYVDYAQW-QRDWMAAGEKQRQLDYWRTRLGTEHPVLELPLDHPRPALQSHRGARRQV 2803
            P +Y DY  W QR  +          +W+  L        +P D  RP    H G+   V
Sbjct: 3410 PPRYRDYIGWLQRQGLDDARA-----WWQANLAGFERATAIPSD--RPLRHDHAGSGMVV 3462

Query: 2804 -----QLDNALVAELKALAQRQEVTLFVLLLASFQTLLHRYSGQSRIRVGVPIANR--NR 2856
                 +L+      L+ LAQ  ++T+     A++  +L RYSG   +  GV +A R  + 
Sbjct: 3463 GDCYTRLEMNDGVRLRELAQAHQLTVNTFAQAAWALVLSRYSGDRDVAFGVTVAGRPVSM 3522

Query: 2857 LETERLLGFFVNTQVLQADVHG---QMAFDQLLAQVKQQALDAQAHQDLPFEQLVEALQP 2913
             + +R +G F+N+  L+  +     + +  Q L  + Q  ++ + ++ LP   +    + 
Sbjct: 3523 PQMQRTVGLFINSVALRVQLPAASERCSVRQWLQGLLQSNMELREYEYLPLVAIQACSEL 3582

Query: 2914 ERSLSHNPLFQVMFNHQDNLRAAPLQLPGLD----LQAVDWAGHS-TQFDLNLETEESAE 2968
             +     PLF  +F  ++    AP++   L+    L A   +G + T F L        +
Sbjct: 3583 PKG---QPLFDSLFVFEN----APVETAVLNHAQHLNASSDSGRTHTNFPLTAVCYPG-D 3634

Query: 2969 GLWASLTYATDLFDATTVERLAEHWQNLLRAVVNDASQAVDELAMLSAPQWQQMVEAWND 3028
             L   L++    FD  TVERL   ++ LL A+V      V EL +LSA +   ++E  N 
Sbjct: 3635 DLGLHLSFDQRYFDYPTVERLLAEFKRLLLALVQGFEGEVSELPLLSAQEQGFLLEDCNR 3694

Query: 3029 TSVDYPRERCVHTLFEARALAQPDALALQFKGQALSYGELNRRANRLAHRLIAAGVGPDV 3088
            T   YP E+     FEA+ +A P     +    +  Y  LN  ANRL H L+AAGV  D 
Sbjct: 3695 TERVYPLEQSYIAQFEAQVVAHPQRTVARCLEASYDYAGLNLAANRLGHALVAAGVCVDQ 3754

Query: 3089 LVAVHVERSLDMVIGLLATLKAGGAYVPLDPQFPADRLAFMLEDSRARVLLTQS------ 3142
             VA+  ER L ++  ++ + KAG  Y+PLDP  PA RL  +++ SR  VL+  +      
Sbjct: 3755 PVALLAERGLPLLGMIVGSFKAGAGYLPLDPGLPAARLQSIIQLSRTPVLVCSAACAEQG 3814

Query: 3143 -HLDGSLAHPQGVQVLM---VEADDNAVEHDPQVNVTPEHLAYVIYTSGSTGTPKGVMVR 3198
              L G L      Q+L+   V+A+  A  H+P ++  P++LAYVIYTSGSTG PKGVMV 
Sbjct: 3815 RQLLGELEASLRPQLLVWEDVQANPIA-SHNPGIHSGPDNLAYVIYTSGSTGLPKGVMVE 3873

Query: 3199 HEALCSFTCGMAGTLDIGADARVLSLTTFSFDIFALELYVPLTVGATVLLSGQELALDPE 3258
               + +        L +     +    + SFDI   +       GA V +    +A DP+
Sbjct: 3874 QRGMLNNQLSKVPYLALSEQDVIAQTASQSFDISVWQFLAAPLFGAKVEIVPNVIAHDPQ 3933

Query: 3259 AIIDLVHSQCANVLQATPSTWRMLLESPRAQLLRGIK-CLCGGEALPSDLAQRLLDL--Q 3315
             ++  V +    VL++ PS  + +L S   Q L G++  L  GE +P +LA + L    Q
Sbjct: 3934 GLLAHVQATGITVLESVPSLIQGMLASDH-QALDGLRWMLPTGEVMPPELAAQWLQRYPQ 3992

Query: 3316 GPLWNLYGPTETTIWSAAHRLHEAQPL-----VGRPIANTALFILNAGLTPSPVGAAGEL 3370
              L N YGP E +   A  R+  A        +G P  N  L++        P+GA GEL
Sbjct: 3993 IGLVNAYGPAECSDDVAFFRVDAASTQGSYLPIGTPTDNNRLYLFGEDQALVPLGAVGEL 4052

Query: 3371 LIGGVGLARGYHERPALTAERFVPNPYGAPGERLYRTGDLARYRADGVVEYIGRVDHQVK 3430
             + G G+ RGY   P  TA  F+P+PYGAPGERLYRTGDLAR R DGV+EY+GR+DHQVK
Sbjct: 4053 CVAGAGVGRGYVGDPVRTALAFIPHPYGAPGERLYRTGDLARQRLDGVLEYVGRIDHQVK 4112

Query: 3431 VRGFRIELGEVEACLREQAGIREAVVL----ADNDRLIAYLVSS---APDAP--EVYKAA 3481
            +RG+RIELGE+EA L EQA IR+A V      +   L+ YLV+    + DA   E  K  
Sbjct: 4113 IRGYRIELGEIEARLHEQAEIRDAAVAVQEGVNGKHLVGYLVAHQGVSADAALLEQVKQR 4172

Query: 3482 LRERLPDYMVPAHLMFLDSLPLTPNGKLDRKALPAVD-AALLRTGHVAPVTPREQQVAAI 3540
            LR  LP+YMVP H  + DSLP   NGKLDRKALPA+D        ++AP    E  +A I
Sbjct: 4173 LRAELPEYMVPLHWGWFDSLPHNANGKLDRKALPAIDIGGQYSQAYLAPRNELEDVLAGI 4232

Query: 3541 WADVLDLPQVGLDDHFFELGGHSLLATRVVSRVRQALALEVALKTLFE---HPLLGDFVQ 3597
            WADVL   +VG+ D+FFELGGHSLLAT++ SRV++ L L V L+ +FE      L ++VQ
Sbjct: 4233 WADVLKAERVGVHDNFFELGGHSLLATQIASRVQKQLQLNVPLRAMFECSTVEALAEYVQ 4292

Query: 3598 AL 3599
             L
Sbjct: 4293 GL 4294



 Score =  469 bits (1206), Expect = e-134
 Identities = 355/1118 (31%), Positives = 541/1118 (48%), Gaps = 61/1118 (5%)

Query: 3599 LGEEGVVAPALLKADRRQPLPLSYAQERQWFLWQLDPHSTAYHIPSALRLHGQLDLGALQ 3658
            L +E + A  +  A      PL+  QE       L+P +  Y++    R++  LD     
Sbjct: 3224 LTQEQIDALPVPAAQIEDVYPLTPMQEGMLLHTLLEPGTGLYYMQDRYRINSALDPERFA 3283

Query: 3659 RSFDTLLARHESLRTHVRQDAA-ATVQVIEDSGLIEIDLAD-------TDEASLKARVAE 3710
            +++  ++ARHE+LR     ++  A +Q+I   G + +D  D         E  L+A   +
Sbjct: 3284 QAWQAVVARHEALRASFSWNSGEAMLQIIHKPGNLAVDYQDWRGLADDAQEQRLQALHKQ 3343

Query: 3711 VVAQPFELLRGPLLRVKLLRLAADDHVLVLVQHHIVSDGWSMQLMVEELVQLYAAFSQGH 3770
                 F LL      ++L+R+A + +  ++  HHI+ D W   L++ +  ++Y A  +G 
Sbjct: 3344 EREAGFALLSEAPFHLRLVRVAEERYWFMMSNHHILIDAWCRSLLMNDFFEVYQALGEGR 3403

Query: 3771 MPDLPALPIQYADYALW-QRNWMEAGEKARQLAYWQAQLGGVQPVLELPFDYLRPAVQSH 3829
               LP +P +Y DY  W QR  ++        A+WQA L G +    +P D  RP    H
Sbjct: 3404 QAQLP-VPPRYRDYIGWLQRQGLDDAR-----AWWQANLAGFERATAIPSD--RPLRHDH 3455

Query: 3830 RGARLGI-----ELQPQLLAGLRRLAQGAGVTLPMVLLASYQALLHRYSGQEDVRVGVPI 3884
             G+ + +      L+      LR LAQ   +T+     A++  +L RYSG  DV  GV +
Sbjct: 3456 AGSGMVVGDCYTRLEMNDGVRLRELAQAHQLTVNTFAQAAWALVLSRYSGDRDVAFGVTV 3515

Query: 3885 ANR--NRLETEGLIGFFVNTQVLKADIHG---QMSVVQLLQQVRQRSLEAQAHQDLPFEQ 3939
            A R  +  + +  +G F+N+  L+  +     + SV Q LQ + Q ++E + ++ LP   
Sbjct: 3516 AGRPVSMPQMQRTVGLFINSVALRVQLPAASERCSVRQWLQGLLQSNMELREYEYLPLVA 3575

Query: 3940 LVEALQPERSMSLSPLFQVLF---NHRV-TSAAHHLQRLTELDVEVLSWDEGVAQFDLAL 3995
            +    +  +     PLF  LF   N  V T+  +H Q L        S D G    +  L
Sbjct: 3576 IQACSELPKGQ---PLFDSLFVFENAPVETAVLNHAQHLNA------SSDSGRTHTNFPL 3626

Query: 3996 D-VEESQATLRASLNYATDLFAPATIERMAGHWQNLLQAMVADQQQPISQLNLLGQDEQH 4054
              V      L   L++    F   T+ER+   ++ LL A+V   +  +S+L LL   EQ 
Sbjct: 3627 TAVCYPGDDLGLHLSFDQRYFDYPTVERLLAEFKRLLLALVQGFEGEVSELPLLSAQEQG 3686

Query: 4055 HILQLWDQTDAGFSAKRLVHELVADRARENPDAVAVKFDAQTLSYGELDRQANRLAHALI 4114
             +L+  ++T+  +  ++        +   +P     +    +  Y  L+  ANRL HAL+
Sbjct: 3687 FLLEDCNRTERVYPLEQSYIAQFEAQVVAHPQRTVARCLEASYDYAGLNLAANRLGHALV 3746

Query: 4115 ARGVGPDVRVAIAMPRSAEIMVAFLAVMKAGGVYVPLDIEYPRDRLLYMMQDSRAQLLLT 4174
            A GV  D  VA+   R   ++   +   KAG  Y+PLD   P  RL  ++Q SR  +L+ 
Sbjct: 3747 AAGVCVDQPVALLAERGLPLLGMIVGSFKAGAGYLPLDPGLPAARLQSIIQLSRTPVLVC 3806

Query: 4175 HTRALQQ-LPIPDGLDSLAIDRTEEWAG-----FSDTAPQVKLGGDNLAYVIYTSGSTGM 4228
                 +Q   +   L++    +   W        +   P +  G DNLAYVIYTSGSTG+
Sbjct: 3807 SAACAEQGRQLLGELEASLRPQLLVWEDVQANPIASHNPGIHSGPDNLAYVIYTSGSTGL 3866

Query: 4229 PKGVAVSHGPLVAHIIATGERYETSPADCELHFMSFAFDGSHEGWMHPLINGASVLIRDD 4288
            PKGV V    ++ + ++       S  D      S +FD S   ++   + GA V I  +
Sbjct: 3867 PKGVMVEQRGMLNNQLSKVPYLALSEQDVIAQTASQSFDISVWQFLAAPLFGAKVEIVPN 3926

Query: 4289 SLWL-PEYTYQQMHRHHVTMAVFPPVYLQQL--AEHAERDGNPPKVRVYCFGGDAVAQAS 4345
             +   P+     +    +T+    P  +Q +  ++H   DG    +R     G+ V    
Sbjct: 3927 VIAHDPQGLLAHVQATGITVLESVPSLIQGMLASDHQALDG----LRWMLPTGE-VMPPE 3981

Query: 4346 YDLAWRALKPTY-LFNGYGPTETVVTPLLWKARKGDPCGAVYAPIGTLLGNRSGYVLDAQ 4404
                W    P   L N YGP E       ++       G+ Y PIGT   N   Y+    
Sbjct: 3982 LAAQWLQRYPQIGLVNAYGPAECSDDVAFFRVDAASTQGS-YLPIGTPTDNNRLYLFGED 4040

Query: 4405 LNLQPIGVAGELYLGGEGVARGYLERPALTAERFVPDPFGKPGSRVYRSGDLTRGRPDGG 4464
              L P+G  GEL + G GV RGY+  P  TA  F+P P+G PG R+YR+GDL R R DG 
Sbjct: 4041 QALVPLGAVGELCVAGAGVGRGYVGDPVRTALAFIPHPYGAPGERLYRTGDLARQRLDGV 4100

Query: 4465 VDYLGSVAHQVKIRGFRIELGEIEARLREQDNVGETVVVAHEGPTGKQLVAYVVPADTLL 4524
            ++Y+G + HQVKIRG+RIELGEIEARL EQ  + +  V   EG  GK LV Y+V    + 
Sbjct: 4101 LEYVGRIDHQVKIRGYRIELGEIEARLHEQAEIRDAAVAVQEGVNGKHLVGYLVAHQGVS 4160

Query: 4525 ADETEFRDSLRRALKTHLPDYMVPTHFMFLAQMPLTPNGKLDRKGLPEPD-ASLLQQAYV 4583
            AD     + +++ L+  LP+YMVP H+ +   +P   NGKLDRK LP  D      QAY+
Sbjct: 4161 AD-AALLEQVKQRLRAELPEYMVPLHWGWFDSLPHNANGKLDRKALPAIDIGGQYSQAYL 4219

Query: 4584 APETELEQQIAAIWADVLRLPQVGLNDNFFEVGGHSLLAIQITSRVQAELGLEVPLLEVF 4643
            AP  ELE  +A IWADVL+  +VG++DNFFE+GGHSLLA QI SRVQ +L L VPL  +F
Sbjct: 4220 APRNELEDVLAGIWADVLKAERVGVHDNFFELGGHSLLATQIASRVQKQLQLNVPLRAMF 4279

Query: 4644 QTETLRAYVQAAATFRAGSAED--FDDLRDFLSELEAI 4679
            +  T+ A  +     +  + +D   D L D ++ELE +
Sbjct: 4280 ECSTVEALAEYVQGLQGSALDDDKVDRLSDLMAELEGL 4317



 Score =  203 bits (516), Expect = 1e-54
 Identities = 289/1127 (25%), Positives = 461/1127 (40%), Gaps = 151/1127 (13%)

Query: 505  EAYPTERGIHHLVEDQVQATPDAPALVFGTTT------LTYAQLDARANQLAHALREKGV 558
            +A+     +   +  +   TP+  AL F   T      L+Y  LD RA  +A AL+ +  
Sbjct: 3    DAFELPLSLVQALAQRAAQTPERIALRFLADTPGEQAVLSYRDLDQRARTIAAALQGRAS 62

Query: 559  APDVLVGICIERSIEMVVGLLAILKAGGAYVPLDP-----EYPQERLAYMIEDSGIQLLL 613
              D  V +      + V      L AG   VP  P     +  QERL  +I+D+  +LLL
Sbjct: 63   FGDRAV-LLFPSGPDYVAAFFGCLYAGVIAVPAYPPESARQQHQERLLSIIDDAEPRLLL 121

Query: 614  SQHSLLPVLPVEGIQVIALDQPASWL--DGYS-AESANVSLHAL---NLAYVIYTSGSTG 667
            +  +L   L  +G++ +A D     L  DG   A +AN     L   ++A++ YTSGST 
Sbjct: 122  TVAALRDSL--QGLEALAADNAPQLLAVDGLDPALAANWREPVLKGDDIAFLQYTSGSTA 179

Query: 668  KPKGAGNSHRALVNRLCWMQQAYGLDAR--DAVLQKTPFSFDVS-VWEFFWPLMTGARLV 724
             PKG   SH  LV     ++Q +G+D    D ++   P   D+  +     P+ +G   V
Sbjct: 180  LPKGVQVSHGNLVANEQLIRQGFGIDLNPDDVIVSWLPLYHDMGLIGGLLQPIFSGVPCV 239

Query: 725  VAAPGEH-REPARLIETIGRHGITTLHFVPSMLQAFIHEPGVQAC-----TSLQRIVCSG 778
            + +P      P R ++ I  +G  T+   P        E   +A       S  R+  SG
Sbjct: 240  LMSPRYFLARPLRWLQAISEYG-GTISGGPDFAYRLCSERVSEASLAGLDLSRWRVAYSG 298

Query: 779  EALPLDAQLQVFAKLPQA------GLFNLYGPTEAAIDVT------HWTCVDEGADSVPI 826
                    L  FA   QA        F  YG  EA + V+          ++  A +   
Sbjct: 299  SEPIRQDSLATFADKFQACGFDPQSFFASYGLAEATLFVSGSRRGQGIAALELDAKAFAA 358

Query: 827  GRPIANLGTYVLDA---------------QLNPVPAGVSGELYLGGIGLARSYHRRPALT 871
             R     G+ ++                 QL  +     GE++ GG  +AR Y R P  +
Sbjct: 359  NRAEPGTGSVLMSCGFPQPGHAVRIVEPQQLQVLGDNQVGEIWAGGPSIARGYWRNPEAS 418

Query: 872  AERFVPSPFGDGARLYRTGDRVRQRADGVIEYLGRLDHQVKLRGLRIELGEIETRLMQHP 931
            A  FV     DG    RTGD    R +G +   GRL   + +RG  +   ++E  L +  
Sbjct: 419  ARTFVEM---DGQTWLRTGDLGFIR-EGEVFVTGRLKDMLIVRGQNLYPQDLEKALEREV 474

Query: 932  SV----REAVVLVQ-------GGKQLVAYLVLEDTAPVDLKAWLLSSLPEYMVPSHMVPL 980
             V    R AV  V+       G    ++  V +   P DL   L   + +    +  V L
Sbjct: 475  QVLRKGRVAVFAVEHQGEEGIGVAVEISRNVQKALKPQDLIKTLRQVVADACRQAPAVVL 534

Query: 981  ----AKLPVTANGKLDRKALPL-------------PDVAPQQAFVAPENDLQKAVAAIWS 1023
                  LP T++GKL R A  L             P+V+         ++LQ  +AA+W 
Sbjct: 535  LLNPGALPKTSSGKLQRSACRLRMDDGSLDCYARFPEVSAAGIRATAGDELQARIAAVWR 594

Query: 1024 DVLGVEQVGLDDNFFELGGDSIISIQVVSR-ARQAGIRLSPRDLFQYQSVRSLALVATFE 1082
            D+L VE V  DD+F  LGG+SI + Q  +R A + GI LS R LF+        ++A + 
Sbjct: 595  DILKVEAVAADDHFLLLGGNSIAATQATARLADELGIDLSLRTLFE------APVLADYS 648

Query: 1083 QATA--IDQGPVTGEVILTPVQHSFFEQAIPARQHW------------NQSLLLTPREAL 1128
            +A A  I  G      I T  +     Q++   + W            N    L  R  L
Sbjct: 649  KAVAQIIADGAAATTAITTLERAQALPQSLAQNRLWLLWQLEPQSAAYNIPAGLHLRGEL 708

Query: 1129 EPARLEAALTRLINHHDALRLRFLQQADGWRQAHAEPVATASLWQSQV---ADEAELAAL 1185
            +   L  A   L+  H++LR  F ++ +G       P    SL +  +   + E   A  
Sbjct: 709  DVNALHVAFQALVARHESLRTVFSEE-NGQALQRILPQQPLSLHRLDLEGHSPEQVAAQR 767

Query: 1186 CDEAQRSLDLEQGPLLRAVLAVMADGTQRLLLVVHHLAVDGVSWRILLEDLQQAY----- 1240
              EA++  DL QGPLLR  L  + D   +L + +HH+  DG S  ILLE+  + Y     
Sbjct: 768  ETEARQPFDLTQGPLLRVTLVRLGDEEHQLWVTLHHIVADGWSLNILLEEFAKLYAARCQ 827

Query: 1241 ----RNAALPAKTSAYQLWAQQLQRHAQTLDEQLPYWQA----QTASADLPCDNPQGGLQ 1292
                  A LP   + Y  W +Q     +  + QL YW+A    +  + DL  D+P+   +
Sbjct: 828  GLEANLAPLPLGYADYGSWQRQWLADGEA-ERQLQYWKAHLGGELPALDLCTDHPRASQR 886

Query: 1293 NRLGSLLEIRLSAEHTRQLLQDAPAAYRTQVNDLLLTALARVISRWSEQPAALIQLEGHG 1352
                +   +++      + L+      +  +  +LL     ++ R+S Q    + +    
Sbjct: 887  EHSAARFSLKVPV-RLGEALKGLAHQQQASLFMVLLAGWQALLQRYSGQADIRVGVPNAN 945

Query: 1353 REDLFDSVDLSRTVGWFTSLFPVRLHADG--ELSAAIKSVKE-QLRAVPHKGIGYGLLRY 1409
            R  L    +    VG+F +   +R   DG    +  +  V++  L+A  ++ + +  L  
Sbjct: 946  RPRL----ETQGMVGFFINTQVLRARFDGRHSFTQLLTQVRQATLQAQANQDLPFEQL-V 1000

Query: 1410 LGTPDAREALSALAAPRITFNYLGQFDRQFNESAL-FVPATQGSGQAQDPQAPLANWLTV 1468
               PDARE        ++ FN+     +Q + SAL  +P       A   +    +    
Sbjct: 1001 EALPDAREQ----GLFQVMFNH-----QQRDLSALRRLPGLLAEELAWHSREAKFDLQLH 1051

Query: 1469 EGQVYGGELALQWGFSREMFDVATVQRLADDYRAELQALIEHCCATP 1515
              + + G L+L + ++ E+F+ +TV+RLA      L AL+E  CA P
Sbjct: 1052 SEEDHLGRLSLAFDYAAELFEASTVERLAH----HLLALLEQVCAAP 1094



 Score =  162 bits (411), Expect = 2e-42
 Identities = 186/683 (27%), Positives = 294/683 (43%), Gaps = 107/683 (15%)

Query: 4063 TDAGFSAKRLVHELVADRARENPDAVAVKFDAQT------LSYGELDRQANRLAHALIAR 4116
            TDA F     + + +A RA + P+ +A++F A T      LSY +LD++A  +A AL  R
Sbjct: 2    TDA-FELPLSLVQALAQRAAQTPERIALRFLADTPGEQAVLSYRDLDQRARTIAAALQGR 60

Query: 4117 GVGPDVRVAIAMPRSAEIMVAFLAVMKAGGVYVPLDIEYP--------RDRLLYMMQDSR 4168
                D R  +  P   + + AF   + AG + VP    YP        ++RLL ++ D+ 
Sbjct: 61   ASFGD-RAVLLFPSGPDYVAAFFGCLYAGVIAVPA---YPPESARQQHQERLLSIIDDAE 116

Query: 4169 AQLLLTHTRALQQLPIPDGLDSLAIDRTEEWAGFSDTAPQVK-------LGGDNLAYVIY 4221
             +LLLT       L    GL++LA D   +        P +        L GD++A++ Y
Sbjct: 117  PRLLLTVAALRDSL---QGLEALAADNAPQLLAVDGLDPALAANWREPVLKGDDIAFLQY 173

Query: 4222 TSGSTGMPKGVAVSHGPLVAH--IIATGERYETSPADCELHFMSFAFD-GSHEGWMHPLI 4278
            TSGST +PKGV VSHG LVA+  +I  G   + +P D  + ++    D G   G + P+ 
Sbjct: 174  TSGSTALPKGVQVSHGNLVANEQLIRQGFGIDLNPDDVIVSWLPLYHDMGLIGGLLQPIF 233

Query: 4279 NGASVLIRDDSLWL--PEYTYQQMHRHHVTMAVFPPVYLQQLAEHAERDG----NPPKVR 4332
            +G   ++     +L  P    Q +  +  T++  P    +  +E          +  + R
Sbjct: 234  SGVPCVLMSPRYFLARPLRWLQAISEYGGTISGGPDFAYRLCSERVSEASLAGLDLSRWR 293

Query: 4333 VYCFGGDAVAQ---------------------ASYDLAWRALKPTYLFNGYGPTETVVTP 4371
            V   G + + Q                     ASY LA   L  +    G G     +  
Sbjct: 294  VAYSGSEPIRQDSLATFADKFQACGFDPQSFFASYGLAEATLFVSGSRRGQGIAALELDA 353

Query: 4372 LLWKARKGDP-CGAVYAPIGTLLGNRSGYVLDAQLNLQPIG--VAGELYLGGEGVARGYL 4428
              + A + +P  G+V    G      +  +++ Q  LQ +G    GE++ GG  +ARGY 
Sbjct: 354  KAFAANRAEPGTGSVLMSCGFPQPGHAVRIVEPQ-QLQVLGDNQVGEIWAGGPSIARGYW 412

Query: 4429 ERPALTAERFVPDPFGKPGSRVYRSGDLTRGRPDGGVDYLGSVAHQVKIRGFRIELGEIE 4488
              P  +A  FV       G    R+GDL   R +G V   G +   + +RG  +   ++E
Sbjct: 413  RNPEASARTFVE----MDGQTWLRTGDLGFIR-EGEVFVTGRLKDMLIVRGQNLYPQDLE 467

Query: 4489 ARLREQDNV---GETVVVA--HEGPTG--------KQLVAYVVPADTLLADETEFRDSLR 4535
              L  +  V   G   V A  H+G  G        + +   + P D +        D+ R
Sbjct: 468  KALEREVQVLRKGRVAVFAVEHQGEEGIGVAVEISRNVQKALKPQDLIKTLRQVVADACR 527

Query: 4536 RALKTHLPDYMVPTHFMFL--AQMPLTPNGKLDRKG--LPEPDASLLQQAYVAPET---- 4587
            +A          P   + L    +P T +GKL R    L   D SL   A   PE     
Sbjct: 528  QA----------PAVVLLLNPGALPKTSSGKLQRSACRLRMDDGSLDCYARF-PEVSAAG 576

Query: 4588 -------ELEQQIAAIWADVLRLPQVGLNDNFFEVGGHSLLAIQITSRVQAELGLEVPLL 4640
                   EL+ +IAA+W D+L++  V  +D+F  +GG+S+ A Q T+R+  ELG+++ L 
Sbjct: 577  IRATAGDELQARIAAVWRDILKVEAVAADDHFLLLGGNSIAATQATARLADELGIDLSLR 636

Query: 4641 EVFQTETLRAYVQAAATFRAGSA 4663
             +F+   L  Y +A A   A  A
Sbjct: 637  TLFEAPVLADYSKAVAQIIADGA 659